Erythranthe tilingii |
Erythranthe filicaulis |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
Hetch Hetchy monkey-flower, slender-stem monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Annuals, taprooted. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
erect, simple or branched at basal nodes, 4–30 cm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
cauline, basal usually not persistent; petiole 0 mm; blade 1-veined or palmately 3-veined, linear to oblanceolate, 6–15 × (1–)2–3 mm, base attenuate, margins entire to remotely mucronulate, teeth 1–3 per side, apex acute, surfaces densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, 2–8, from medial to distal nodes. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
pink to purple, throat dark red-purple, palate ridges yellow, bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat cylindric-funnelform, 8–12(–17) mm, exserted beyond calyx margin; limb expanded 10–14 mm, lobes notched, palate villous. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
9–30 mm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
reddish brown-spotted, strongly angled, cylindric-campanulate, 7–9 mm, margins distinctly toothed or lobed, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped, ribs corky, lobes pronounced, spreading. |
Capsules | included, 5–7 mm. |
included, 5–7 mm. |
Anthers | included, glabrous. |
included, hirtellous. |
2n | = 28, 56. |
= 16. |
Erythranthe tilingii |
Erythranthe filicaulis |
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Phenology | Flowering Jul–Sep. | Flowering May–Jul. |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Moist open areas on gentle slopes, meadows, roadsides, gravelly soils. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 800–1800 m. (2600–5900 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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CA
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Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe filicaulis is known only for certain from Mariposa and Tuolumne counties. Kappler 1691 (UCLA) from Madera County also may be this species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 391. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | Mimulus filicaulis, M. biolettii |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (S. Watson) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) |
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