Erythranthe tilingii |
Erythranthe cardinalis |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
cardinal monkey flower, scarlet monkey-flower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Perennials, rhizomatous. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
usually erect to ascending, freely branched, 25–75 cm, ± glandular-villous or glabrate. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
usually cauline; petiole 0 mm; blade palmately veined, elliptic-ovate to ovate or obovate, 20–90(–110) × 10–38(–60) mm, thick, base acuminate, subclasping, margins irregularly serrate to dentate, apex acute, surfaces ± glandular-villous to glabrate. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, 2–12, axillary at leafy medial to distal nodes. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
scarlet to orange-red, rarely yellow, throat yellowish, red-striped, palate red, yellow-villous, not spotted or striped, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, (15–)20–30 mm, exserted beyond calyx margin; throat open. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
30–90(–120) mm. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
cylindric to cylindric-campanulate, not inflated, 17–28(–30) mm, hispid-hirsute to hirsute, lobes 4–7 mm, ovate to ovate-deltate, apex attenuate-acute. |
Capsules | included, 5–7 mm. |
included, 10–16 mm. |
Anthers | included, glabrous. |
exserted, white-villous, thecae spreading. |
2n | = 28, 56. |
= 16. |
Erythranthe tilingii |
Erythranthe cardinalis |
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Phenology | Flowering Jul–Sep. | Flowering May–Sep. |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Moist to wet places along streams, lakes, creek beds, canyon, arroyo, and ravine bottoms, around springs and seepage areas, flood plains, moist clearings and woods edges. |
Elevation | 1400–3400 m. (4600–11200 ft.) | (5–)50–2300(–2800) m. ((0–)200–7500(–9200) ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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CA; NV; OR; WA; Mexico (Baja California)
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Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Yellow-flowered populations of Erythranthe cardinalis occur on Cedros Island, Baja California, and in the Siskiyou Mountains of Oregon (R. K. Vickery 1992). They have been documented also on Santa Cruz Island, California. Populations of Erythranthe cardinalis in King and Yakima counties, Washington, are introduced and naturalized. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 393. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | Mimulus cardinalis, M. cardinalis var. exsul, M. cardinalis var. griseus, M. cardinalis var. rigens |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Douglas ex Bentham) Spach: Hist. Nat. Vég. 9: 313. (1840) |
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