Erythranthe tilingii |
Erythranthe calciphila |
|
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
Mexican moss monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Annuals, usually fibrous-rooted, rarely rooting at proximal nodes. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
erect, sometimes decumbent-ascending, branched, 4–30 cm, minutely stipitate-glandular, also delicately villosulous-glandular along whole length. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
basal and cauline; petiole 1–3 mm, cauline blade slightly or hardly reduced in size from basal, becoming subsessile to sessile (1–3 pairs of cauline leaves); blade palmately 3(–5)-veined, orbicular-ovate to oblong-ovate, 7–28 × 5–22 mm, base truncate to shallowly cuneate, margins shallowly dentate to denticulate, teeth 3–6 per side, apex rounded to obtuse, surfaces villous, hairs thin-walled, vitreous, eglandular or minutely gland-tipped, usually also minutely stipitate-glandular. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
plesiogamous, 1–6(–10), axillary at all nodes, chasmogamous. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
light yellow, red-spotted, weakly bilaterally symmetric, ± bilabiate; tube-throat narrowly funnelform, 5–7 mm, exserted 1–2 mm beyond calyx margin; limb expanded 2–4 mm. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
15–30(–55) mm, minutely stipitate-glandular, sometimes minutely hirtellous and minutely stipitate-glandular, sometimes short glandular-villous. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
nodding 90º at maturity, usually 3(–5)-veined, ovoid, inflated, sagittally compressed, 6–10 mm, minutely stipitate-glandular, throat closing, lobes usually 3 or 3 and 5 on same plant, if 5 then with 2 interpolated lobes much smaller than abaxial pair. |
Capsules | included, 5–7 mm. |
included, 2–4 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28, 56. |
= 30, 32. |
Erythranthe tilingii |
Erythranthe calciphila |
|
Phenology | Flowering Jul–Sep. | Flowering Mar–Sep(–Nov). |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Rocky knobs, moist boulders, wet rock faces, roadcuts, seepages, springs, with moss, usually in pine or pine-oak woods. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 1800–2500 m. (5900–8200 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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AZ; Mexico (Chihuahua, Durango, Sinaloa, Sonora) |
Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe calciphila is recognized by its annual duration (fibrous-rooted), short, erect stems with few, even-sized leaves (the basal often persistent), delicate stipitate-glandular vestiture, three-lobed calyces that are relatively large in fruit, small corollas, and autogamous reproduction. Plants rarely root at proximal nodes. In Arizona, at the northernmost extremity of its range, E. calciphila occurs in the Chiricahua, Huachuca, and Mule mountains of Cochise County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 424. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | Mimulus calciphilus, M. minutiflorus |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Gentry) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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