Erythranthe tilingii |
Erythranthe calcicola |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
limestone monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Annuals, taprooted. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
erect, simple or branched from basal nodes, 2–10(–15) cm, sparsely glandular-pubescent, internodes elongate, distinct. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
cauline, basal not persistent; petiole 0–1 mm; blade palmately 3-veined (in broader ones), lanceolate to ovate, 3–25 × 2–8(–10) mm, base attenuate, margins entire, sometimes toothed, apex acute to obtuse, surfaces sparsely glandular-pubescent. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, sometimes plesiogamous, 1–16, from distal or medial to distal nodes. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
yellow or white and throat yellow, throat and limb red-spotted, bilaterally symmetric, weakly bilabiate; tube-throat cylindric, 6–13 mm, exserted beyond calyx margin; limb expanded 3–7(–9) mm, lobes notched, abaxial limb sparsely bearded or glabrous. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
(3–)5–20 mm. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
campanulate to widely cylindric, 5–7 mm, margins distinctly toothed or lobed, sparsely glandular-pubescent, ribs strongly angled, lobes pronounced, erect. |
Capsules | included, 5–7 mm. |
included, 4–6 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28, 56. |
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Erythranthe tilingii |
Erythranthe calcicola |
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Phenology | Flowering Jul–Sep. | Flowering Apr–Jun. |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Creosote bush and Joshua tree woodlands, usually on carbonate substrate, primarily on talus slopes. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 900–2200 m. (3000–7200 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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CA; NV |
Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe calcicola was previously included in the broader concept of E. montioides; the former can be distinguished by its glandular herbage, broader lanceolate to ovate leaves, strongly angled calyces, and smaller flowers. The species is restricted to the northern Mojave Desert and southwestern Great Basin and occurs primarily on talus slopes on substrates derived from carbonate rock. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 383. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | N. S. Fraga & D. A. York: Aliso 30: 54, figs. 12–16. (2012) |
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