Erythranthe tilingii |
Erythranthe caespitosa |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
large mountain monkey-flower, mountain monkeyflower, Olympic monkeyflower, subalpine monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Perennials, rhizomatous, rooting at proximal nodes, sometimes producing creeping, small-leaved runners, forming matted colonies, rhizomes filiform. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
procumbent or decumbent to decumbent-ascending, delicate, usually in masses, terete or flattish, branched, 3–10 cm, glabrous, minutely hirtellous, or stipitate-glandular. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
basal and cauline; petiole: proximals 2–5 mm, distals 0 mm; blade often purple beneath, palmately 3-veined, orbicular to narrowly elliptic or ovate, proximals usually sublyrate, 3–12 mm, becoming larger distally, base cuneate to a short petiole, margins entire, mucronulate, or barely denticulate, apex obtuse, surfaces sparsely to moderately puberulent, hairs minute, stipitate-glandular. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, 1–3, from distal nodes, commonly solitary. |
Styles | hirtellous. |
minutely hirtellous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
yellow, dark red-spotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform to cylindric-funnelform, 15–18 mm, exserted beyond calyx margin; abaxial limb with deflexed-spreading lobes, adaxial with ascending lobes, palate partially closed. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
10–30(–40) mm, sparsely to moderately villous, hairs short, gland-tipped, sometimes hirtellous. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
broadly campanulate, inflated, sagittally compressed, 7–15 mm, glabrous, minutely hirtellous, or stipitate-glandular, throat closing, proximalmost lobe pair upcurving, distalmost 3–5 mm, prominently protruding. |
Capsules | included, 5–7 mm. |
included, 4–5 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28, 56. |
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Erythranthe tilingii |
Erythranthe caespitosa |
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Phenology | Flowering Jul–Sep. | Flowering Jul–Sep. |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Alpine meadows and slopes, stream banks, wet rocks in streams, wet crevices, talus. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 1100–2000(–2300) m. (3600–6600(–7500) ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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WA; BC
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Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe caespitosa is endemic to northwestern and central Washington (Cascade and Olympic mountains) and adjacent British Columbia (Cascades, Selkirk Mountains and Chilliwack Valley, Coast Mountains). The plants have consistently small leaves with subentire margins, and the stems are consistently procumbent to decumbent-ascending, usually forming matted colonies. Erythranthe caespitosa and E. tilingii appear to be sympatric in counties of northwestern Washington, but this needs to be verified in the field. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 409. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | Mimulus scouleri var. caespitosus, M. caespitosus, M. tilingii var. caespitosus |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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