Erythranthe tilingii |
Erythranthe acutidens |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
Kings River monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Annuals, fibrous-rooted. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
erect to ascending-erect, simple or diffusely branched, 4-angled, 7–20 cm, glabrous. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
basal and cauline; petiole 0 mm; blade palmately 3–5 veined, ovate to broadly ovate, 10–20 × 7–11 mm, base rounded to truncate, margins serrate-denticulate, apex acute, surfaces glabrous. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, 2–20, from proximal to distal nodes. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
pale pink to rose purple, throat yellow or deep pink externally with 2 yellow spots below abaxial limb, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 9–12 mm, exserted beyond calyx margin; limb expanded 9–12 mm. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
divergent-arcuate, sometimes becoming deflexed, 10–23 mm, longer than subtending leaves, glabrous. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
campanulate, 7–9 mm, margins subtruncate, glabrous, lobes reduced, subequal. |
Capsules | included, 5–7 mm. |
included, (4–)5–10 mm. |
Anthers | included, glabrous. |
included, minutely villous-hirsute. |
2n | = 28, 56. |
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Erythranthe tilingii |
Erythranthe acutidens |
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Phenology | Flowering Jul–Sep. | Flowering Apr–Jul. |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Grassy slopes, sandy terraces, marshy places, lakeshores, creek sides, seep edges, shaded canyon slopes, roadcuts and roadsides, woodlands of Pseudotsuga menziesii-Quercus chrysolepis, oak-gray pine, or interior live oak. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 200–2000 m. (700–6600 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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CA
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Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Mimulus acutidens Greene is heterotypic with the South American M. acutidens Reiche, which is a later homonym. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 383. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | Mimulus acutidens, M. inconspicuus var. acutidens |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 34. (2012) |
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