Erythranthe taylorii |
Erythranthe verbenacea |
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Shasta limestone monkeyflower, Taylor's or Shasta limestone monkeyflower |
crimson monkeyflower, pico Pajaro |
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Habit | Annuals, filiform-taprooted. | Perennials, rhizomatous. |
Stems | erect, straight at nodes, simple or few-branched from base, 5–10 cm, sparsely eglandular-villous proximally, becoming sparsely short stipitate-glandular distally. |
erect to decumbent, usually simple, weakly 4-angled, 20–60 cm, ± glandular-villous. |
Leaves | usually cauline, basal not persistent, largest at mid stem or basal and mid stem to nearly even-sized; petiole 3–5(–8) mm; blade often purple adaxially, palmately 3–5-veined, broadly ovate to elliptic-ovate or ovate-lanceolate, 4–20 × 4–12 mm, base rounded to truncate, margins serrate-dentate, teeth 2–4 per side, shallow, apex rounded to obtuse or acute, surfaces: distals moderately short-stipitate-glandular. |
cauline; petiole 0 mm; blade palmately 3–5-veined, elliptic to obovate, rhombic-ovate, or broadly spatulate, 50–75 × 15–26(–30) mm, base subcordate, subclasping, margins coarsely serrate, sometimes only distally, apex acute to obtuse, surfaces ± glandular-villous. |
Flowers | herkogamous, sometimes plesiogamous, 2–6(–8), from proximal to distal nodes. |
herkogamous, 2–12, axillary at leafy medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat ceiling sometimes red-spotted or -lined, abaxial limb yellow or with 1 or 2 red splotches, bilaterally symmetric, bilabiate; tube-throat funnelform, 5–7 mm, exserted beyond calyx margin. |
crimson, often yellow-tinged, palate ridges dark red, bilaterally symmetric, strongly bilabiate; tube-throat tubular, 25–35 mm, exserted 13–25 mm beyond calyx margin; abaxial limb spreading, adaxial erect, lobe apex truncate, often emarginate, throat open, palate ridges densely short-villous. |
Fruiting pedicels | divergent to arcuate-divergent, 6–13 mm. |
45–90(–150) mm. |
Fruiting calyces | wing-angled, tubular-campanulate, 4–5 mm, margins distinctly toothed or lobed, densely invested with tiny, waxy-white, eglandular, papillose hairs between angles, lobes pronounced, erect. |
campanulate, weakly inflated, 20–28 mm, sparsely glandular-villosulous to stipitate-glandular, lobes triangular to ovate-triangular, apex linear-triangular. |
Capsules | included, 3–4 mm. |
included, 15–22 mm. |
Anthers | included, glabrous. |
exserted, white-villous, thecae reflexed 45º. |
2n | = 16. |
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Erythranthe taylorii |
Erythranthe verbenacea |
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Phenology | Flowering Feb–May. | Flowering Jun–Sep. |
Habitat | Crevices in limestone cliff faces and outcrops. | Stream edges and beds, flood plains, around seeps and springs, canyon bottoms, moist cliff crevices and ledges. |
Elevation | 900–1100 m. (3000–3600 ft.) | 300–2600 m. (1000–8500 ft.) |
Distribution |
CA
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AZ; UT; Mexico (Baja California, Chihuahua, Durango, Sinaloa, Sonora)
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Discussion | Erythranthe taylorii is known only from the Shasta Lake region of northwestern Shasta County. Its broad, distinctly bilabiate corollas and ovate leaf blades with palmate venation are similar to those of species of the northern group of sect. Mimulosma, the "Columbia River clade" (J. B. Whittall et al. 2006) of Idaho, Montana, Oregon, Washington, and Wyoming, particularly to the Idaho endemic E. ampliata. Erythranthe taylorii is distinct from E. ampliata in its larger, papillose calyces, shorter fruiting pedicels, corollas with shorter tube-throats, and shorter capsules. Considerable corolla color variation exists in E. taylorii in the occurrence and density of red dots and lines on the throat ceiling and larger red splotches on the abaxial limb. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
R. K. Vickery (1992) noted that yellow-flowered morphs of Erythranthe verbenacea occur in a population at Vasey’s Paradise in the Grand Canyon (Coconino County), 32 miles downstream from Lees Ferry. Populations of Erythranthe verbenacea in the Oak Creek Canyon area in southern Coconino County, Arizona, have leaves with a narrow, lateral, undulating, purple stripe across the mid lamina. The coloration is retained even in dried specimens. In Utah, Erythranthe verbenacea is known only from the Zion Canyon area. Molecular (P. M. Beardsley et al. 2003) and morphological (G. L. Nesom 2014b) data indicate that Erythranthe verbenacea is sister to E. eastwoodiae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 398. | FNA vol. 17, p. 394. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus verbenaceus, M. cardinalis var. verbenaceus, M. lugens | |
Name authority | G. L. Nesom: Phytoneuron 2013-43: 6, figs. 5–7. (2013) — (as taylori) | (Greene) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 37. (2012) |
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