Erythranthe taylorii |
Erythranthe ptilota |
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Shasta limestone monkeyflower, Taylor's or Shasta limestone monkeyflower |
musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
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Habit | Annuals, filiform-taprooted. | Perennials, rhizomatous, sometimes rooting at proximal nodes. |
Stems | erect, straight at nodes, simple or few-branched from base, 5–10 cm, sparsely eglandular-villous proximally, becoming sparsely short stipitate-glandular distally. |
prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
Leaves | usually cauline, basal not persistent, largest at mid stem or basal and mid stem to nearly even-sized; petiole 3–5(–8) mm; blade often purple adaxially, palmately 3–5-veined, broadly ovate to elliptic-ovate or ovate-lanceolate, 4–20 × 4–12 mm, base rounded to truncate, margins serrate-dentate, teeth 2–4 per side, shallow, apex rounded to obtuse or acute, surfaces: distals moderately short-stipitate-glandular. |
cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
Flowers | herkogamous, sometimes plesiogamous, 2–6(–8), from proximal to distal nodes. |
herkogamous, 4–10, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat ceiling sometimes red-spotted or -lined, abaxial limb yellow or with 1 or 2 red splotches, bilaterally symmetric, bilabiate; tube-throat funnelform, 5–7 mm, exserted beyond calyx margin. |
yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
Fruiting pedicels | divergent to arcuate-divergent, 6–13 mm. |
(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
Fruiting calyces | wing-angled, tubular-campanulate, 4–5 mm, margins distinctly toothed or lobed, densely invested with tiny, waxy-white, eglandular, papillose hairs between angles, lobes pronounced, erect. |
wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
Capsules | included, 3–4 mm. |
included, 6–8 mm. |
Anthers | included, glabrous. |
included, finely hirtellous to hispidulous. |
Erythranthe taylorii |
Erythranthe ptilota |
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Phenology | Flowering Feb–May. | Flowering (May–)Jun–Aug. |
Habitat | Crevices in limestone cliff faces and outcrops. | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. |
Elevation | 900–1100 m. (3000–3600 ft.) | 0–1000(–1900) m. (0–3300(–6200) ft.) |
Distribution |
CA
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CA; OR; WA; BC
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Discussion | Erythranthe taylorii is known only from the Shasta Lake region of northwestern Shasta County. Its broad, distinctly bilabiate corollas and ovate leaf blades with palmate venation are similar to those of species of the northern group of sect. Mimulosma, the "Columbia River clade" (J. B. Whittall et al. 2006) of Idaho, Montana, Oregon, Washington, and Wyoming, particularly to the Idaho endemic E. ampliata. Erythranthe taylorii is distinct from E. ampliata in its larger, papillose calyces, shorter fruiting pedicels, corollas with shorter tube-throats, and shorter capsules. Considerable corolla color variation exists in E. taylorii in the occurrence and density of red dots and lines on the throat ceiling and larger red splotches on the abaxial limb. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 398. | FNA vol. 17, p. 402. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | |
Name authority | G. L. Nesom: Phytoneuron 2013-43: 6, figs. 5–7. (2013) — (as taylori) | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) |
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