Erythranthe taylorii |
Erythranthe inflatula |
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Shasta limestone monkeyflower, Taylor's or Shasta limestone monkeyflower |
disappearing monkey-flower, ephemeral monkeyflower |
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Habit | Annuals, filiform-taprooted. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | erect, straight at nodes, simple or few-branched from base, 5–10 cm, sparsely eglandular-villous proximally, becoming sparsely short stipitate-glandular distally. |
erect to ascending, straight or geniculate at nodes, simple or branched at proximal and medial nodes, 6–20(–25) cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Leaves | usually cauline, basal not persistent, largest at mid stem or basal and mid stem to nearly even-sized; petiole 3–5(–8) mm; blade often purple adaxially, palmately 3–5-veined, broadly ovate to elliptic-ovate or ovate-lanceolate, 4–20 × 4–12 mm, base rounded to truncate, margins serrate-dentate, teeth 2–4 per side, shallow, apex rounded to obtuse or acute, surfaces: distals moderately short-stipitate-glandular. |
usually cauline, basal usually deciduous by flowering; petiole: proximals 1–3 mm, distals 0 mm; blade palmately 3–5-veined, narrowly ovate or narrowly lanceolate to elliptic or elliptic-lanceolate, largest 8–18(–30) × (1–)3–7 mm, relatively even-sized, or slightly reduced distally, base attenuate to obtuse or rounded, margins entire, mucronulate, or denticulate, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Flowers | herkogamous, sometimes plesiogamous, 2–6(–8), from proximal to distal nodes. |
plesiogamous, 10–20, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat ceiling sometimes red-spotted or -lined, abaxial limb yellow or with 1 or 2 red splotches, bilaterally symmetric, bilabiate; tube-throat funnelform, 5–7 mm, exserted beyond calyx margin. |
yellow to pale yellow, sparsely red-spotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric, 5–8 mm, exserted 1–3 mm beyond calyx margin; limb barely widened, lobes broadly obovate, apex rounded or mucronate. |
Fruiting pedicels | divergent to arcuate-divergent, 6–13 mm. |
straight, 7–18 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Fruiting calyces | wing-angled, tubular-campanulate, 4–5 mm, margins distinctly toothed or lobed, densely invested with tiny, waxy-white, eglandular, papillose hairs between angles, lobes pronounced, erect. |
winged, plicate-angled, maturing ovoid-ellipsoid to campanulate or broadly urceolate, distinctly inflated, 7–11 mm, margins distinctly toothed or lobed, sparsely, minutely hirtellous, eglandular, lobes pronounced, erect. |
Capsules | included, 3–4 mm. |
included, 5–9 mm. |
Anthers | included, glabrous. |
included, glabrous. |
Erythranthe taylorii |
Erythranthe inflatula |
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Phenology | Flowering Feb–May. | Flowering Jun–Jul. |
Habitat | Crevices in limestone cliff faces and outcrops. | Drying edges, banks, and beds of summer-dry watercourses, near drying edges of small lakes or impoundments, often among rocks and shoreline detritus, occasionally in moist protected areas beneath low shrubs. |
Elevation | 900–1100 m. (3000–3600 ft.) | 1200–1700 m. (3900–5600 ft.) |
Distribution |
CA
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CA; ID; NV; OR |
Discussion | Erythranthe taylorii is known only from the Shasta Lake region of northwestern Shasta County. Its broad, distinctly bilabiate corollas and ovate leaf blades with palmate venation are similar to those of species of the northern group of sect. Mimulosma, the "Columbia River clade" (J. B. Whittall et al. 2006) of Idaho, Montana, Oregon, Washington, and Wyoming, particularly to the Idaho endemic E. ampliata. Erythranthe taylorii is distinct from E. ampliata in its larger, papillose calyces, shorter fruiting pedicels, corollas with shorter tube-throats, and shorter capsules. Considerable corolla color variation exists in E. taylorii in the occurrence and density of red dots and lines on the throat ceiling and larger red splotches on the abaxial limb. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
No natural occurrences of Erythranthe inflatula are known from Washington; the type collection from Klickitat County is from a cultivated plant. Morphological and molecular data (R. J. Meinke 1995; P. M. Beardsley et al. 2004) indicate that Erythranthe inflatula originated as a hybrid between E. breviflora and E. latidens. Its geography and biology suggest that it is reproductively stable. The putative parents are geographically and ecologically separated for most of their ranges, and the range of E. inflatula is considerably broader than the relatively small region where the parents are sympatric. In the region of sympatry, however, E. inflatula may be difficult to distinguish from one or both of its putative parents. G. L. Nesom (2012g) was not able to find morphology that would distinguish the recently described Mimulus evanescens from E. inflatula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 398. | FNA vol. 17, p. 400. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus inflatulus, M. evanescens | |
Name authority | G. L. Nesom: Phytoneuron 2013-43: 6, figs. 5–7. (2013) — (as taylori) | (Suksdorf) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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