Erythranthe rubella |
Erythranthe ptilota |
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little redstem monkeyflower, red stem himulus, redstem monkeyflower, redstem or little redstem monkeyflower |
musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
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Habit | Annuals, taprooted. | Perennials, rhizomatous, sometimes rooting at proximal nodes. |
Stems | erect, simple, sometimes branched from basal nodes, 3–32 cm, minutely puberulent. |
prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
Leaves | cauline, basal not persistent; petiole 0 mm; blade palmately 3-veined (in broader ones), linear to elliptic, 5–22(–30) × 1–5 mm, base narrowed, margins entire, sometimes toothed, apex acute to obtuse, surfaces minutely puberulent. |
cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
Flowers | herkogamous, sometimes plesiogamous, 1–106, from distal or medial to distal nodes. |
herkogamous, 4–10, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow and abaxial limb and throat red dotted or pink to purple and throat yellow, bilaterally symmetric, weakly bilabiate; tube-throat cylindric, 4–10 mm, exserted beyond calyx margin; limb expanded 3–5 mm, lobes entire or weakly notched, abaxial limb glabrous. |
yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
Fruiting pedicels | 2–18 mm. |
(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
Fruiting calyces | becoming red-angled or red, campanulate to nearly cylindric, 4–9 mm, margins distinctly toothed or lobed, glabrous or minutely puberulent, ribs thickened, lobes pronounced, erect, margins ciliate. |
wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
Capsules | included, 3–8 mm. |
included, 6–8 mm. |
Anthers | included, glabrous. |
included, finely hirtellous to hispidulous. |
Erythranthe rubella |
Erythranthe ptilota |
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Phenology | Flowering Apr–Jun. | Flowering (May–)Jun–Aug. |
Habitat | Open slopes and washes. | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. |
Elevation | 300–3000 m. [1000–9800 ft.] | 0–1000(–1900) m. [0–3300(–6200) ft.] |
Distribution |
AZ; CA; CO; NM; NV; TX; UT; WY; Mexico (Baja California, Sonora)
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CA; OR; WA; BC
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Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 17, p. 384. | FNA vol. 17, p. 402. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Mimulus rubellus, M. gratioloides | Mimulus moschatus var. sessilifolius |
Name authority | (A. Gray) N. S. Fraga: Phytoneuron 2012-39: 35. (2012) | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) |
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