Erythranthe pulsiferae |
Erythranthe tilingii |
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candelabrum monkey-flower, Pulsifer's monkey-flower |
larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
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Habit | Annuals, shallowly fibrous-rooted. | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. |
Stems | erect, straight at nodes, simple or few-branched at base, 5–12(–18) cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
Leaves | basal and cauline; petiole 2–9 mm, distinctly 3-veined, 2-winged; blade palmately 3-veined, elliptic-oblong to ovate or oblanceolate, 3–14(–23) × 2–9(–15) mm, base cuneate to attenuate, margins denticulate to entire, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
Flowers | herkogamous, 1–9, from medial to distal nodes. |
herkogamous, 1–3(–5), from distal nodes. |
Styles | glabrous. |
hirtellous. |
Corollas | yellowish, tube-throat, palate ridges, and limb yellow to pale yellow, abaxial limb red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 6–9 mm, exserted beyond calyx margin; lobes broadly obovate-suborbicular, apex rounded. |
yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
Fruiting pedicels | divergent-arcuate, 12–38 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
Fruiting calyces | cylindric, ± inflated, 7–10 mm, margins distinctly toothed or lobed, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped, lobes pronounced, erect. |
usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
Capsules | included, 5–8 mm. |
included, 5–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 28, 56. |
Erythranthe pulsiferae |
Erythranthe tilingii |
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Phenology | Flowering Apr–Jul. | Flowering Jul–Sep. |
Habitat | Damp depressions, moist gravel, rocky flats, granite outcrops, wet meadows, lava beds, vernal pools, forest openings, commonly in or near coniferous forests, chaparral-live oak woodlands. | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. |
Elevation | 50–1300(–2500) m. (200–4300(–8200) ft.) | 1400–3400 m. (4600–11200 ft.) |
Distribution |
CA; OR; WA
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AZ; CA; ID; MT; NV; OR; UT; AB
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Discussion | Erythranthe pulsiferae is characterized by minutely stipitate-glandular vestiture (lacking villous hairs), elongate internodes, persistent basal leaves, small, palmately veined, cauline leaves with short, three-veined petioles and elliptic-oblong to ovate or oblanceolate blades, divergent-arcuate pedicels, and small, all yellow, weakly bilabiate corollas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 398. | FNA vol. 17, p. 408. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus pulsiferae | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius |
Name authority | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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