Erythranthe pulsiferae |
Erythranthe moschata |
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candelabrum monkey-flower, Pulsifer's monkey-flower |
mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower |
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Habit | Annuals, shallowly fibrous-rooted. | Perennials, rhizomatous, rooting at proximal nodes. |
Stems | erect, straight at nodes, simple or few-branched at base, 5–12(–18) cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident. |
Leaves | basal and cauline; petiole 2–9 mm, distinctly 3-veined, 2-winged; blade palmately 3-veined, elliptic-oblong to ovate or oblanceolate, 3–14(–23) × 2–9(–15) mm, base cuneate to attenuate, margins denticulate to entire, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
usually cauline, basal not persistent, distinctly separated; petiole 0 mm or (0.5–)1–5(–10) mm; blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous. |
Flowers | herkogamous, 1–9, from medial to distal nodes. |
herkogamous, 1–8, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellowish, tube-throat, palate ridges, and limb yellow to pale yellow, abaxial limb red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 6–9 mm, exserted beyond calyx margin; lobes broadly obovate-suborbicular, apex rounded. |
yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin; lobes oblong-obovate, apex rounded to notched. |
Fruiting pedicels | divergent-arcuate, 12–38 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
(7–)10–25 mm, glabrate to glandular-villous. |
Fruiting calyces | cylindric, ± inflated, 7–10 mm, margins distinctly toothed or lobed, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped, lobes pronounced, erect. |
ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse. |
Capsules | included, 5–8 mm. |
included, 6–8 mm. |
Anthers | included, glabrous. |
included, glabrous or slightly hirtellous to scabrous. |
2n | = 32. |
= 32. |
Erythranthe pulsiferae |
Erythranthe moschata |
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Phenology | Flowering Apr–Jul. | Flowering May–Aug. |
Habitat | Damp depressions, moist gravel, rocky flats, granite outcrops, wet meadows, lava beds, vernal pools, forest openings, commonly in or near coniferous forests, chaparral-live oak woodlands. | Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. |
Elevation | 50–1300(–2500) m. (200–4300(–8200) ft.) | (300–)400–3100 m. ((1000–)1300–10200 ft.) |
Distribution |
CA; OR; WA
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CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
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Discussion | Erythranthe pulsiferae is characterized by minutely stipitate-glandular vestiture (lacking villous hairs), elongate internodes, persistent basal leaves, small, palmately veined, cauline leaves with short, three-veined petioles and elliptic-oblong to ovate or oblanceolate blades, divergent-arcuate pedicels, and small, all yellow, weakly bilabiate corollas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size. Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 398. | FNA vol. 17, p. 401. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus pulsiferae | Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus |
Name authority | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) | (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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