Erythranthe ptilota |
Erythranthe unimaculata |
|
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
green-palate monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, shallowly fibrous-rooted or slender-taprooted, sometimes rooting at proximal nodes. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect or basally ascending-erect, simple or few-branched, becoming fistulose in larger plants, (2–)10–30(–100) cm, delicately short glandular-villous to stipitate-glandular, often glabrous below inflorescence. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
usually cauline, basal sometimes persistent, distal connate-perfoliate, often bractlike; petiole: proximals 4–10(–15) mm, distals 0 mm; blade: proximals sometimes subpinnately veined, usually with (1 or)2 pairs arising from midvein above base, becoming palmately veined distally, ovate-lanceolate to ovate or broadly ovate-elliptic, mid cauline 12–40(–50) × 10–25(–45) mm, base rounded to truncate or cuneate, margins shallowly dentate-serrate to serrate, teeth 7–12 per side, apex rounded to obtuse, surfaces glabrous, sometimes with sharp-pointed hirtellous, vitreous-flattened, or gland-tipped hairs. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, (1–)3–14, usually from mid stem and distally, chasmogamous. |
Styles | glabrous. |
hirtellous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
pale yellow, palate and abaxial throat dark yellow, drying blue-green, red-spotted, bilaterally symmetric, bilabiate; tube-throat funnelform to subfunnelform, (7–)9–14 mm, exserted 3–4 mm beyond calyx margin; limb broadly expanded (8–17 mm pressed), palate densely bearded. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
10–40 mm, delicately short glandular-villous to stipitate-glandular, often glabrous below inflorescence. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
nodding 30–90º, broadly campanulate, inflated, sagittally compressed, 9–13(–15) mm, sparsely glandular-villous to stipitate-glandular, throat closing, adaxial lobe distinctly longer. |
Capsules | included, 6–8 mm. |
included, 5–8 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
2n | = 28. |
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Erythranthe ptilota |
Erythranthe unimaculata |
|
Phenology | Flowering (May–)Jun–Aug. | Flowering Jan–Jun. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Stream and canal sides, pool edges, canyon bottoms, sand, gravel, and mud, riparian habitats, pine-oak forests. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 200–2000 m. (700–6600 ft.) |
Distribution |
CA; OR; WA; BC
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AZ; NM; Mexico (Chihuahua, Sonora) |
Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe unimaculata is recognized by its annual duration (fibrous-rooted, without stolons or rhizomes), delicate-glandular vestiture, mostly sessile to subsessile, often widely spaced leaves, closed fruiting calyces, and relatively large, pale yellow to nearly white corollas with a dark yellow palate that commonly dries blue-green. The breeding system is allogamous. Plants commonly are single-stemmed and usually erect but sometimes produce decumbent-ascending stems branching from the base, these sometimes rooting at proximal nodes. Erythranthe unimaculata is known from Cochise, Gila, Pima, and Santa Cruz counties, Arizona, and from Doña Ana County, New Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 413. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus unimaculatus |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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