Erythranthe ptilota |
Erythranthe sierrae |
|
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
Sierra Nevada monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, taprooted. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, simple or branched from basal nodes, (3–)4–20 cm, minutely glandular. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
cauline, basal not persistent; petiole 0 mm; blade 1-veined or palmately 3-veined (in broader ones), linear to oblanceolate, 3–23 × 1–10 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces minutely glandular. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 1–38, from distal or medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
pale pink, abaxial limb with 2 yellow ridges, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 5–15 mm, exserted beyond calyx margin; limb expanded 5–17 mm, lobes deeply notched, abaxial limb glabrous or sparsely puberulent. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
10–33(–40) mm. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
sometimes red-spotted, campanulate to cylindric, 4–8 mm, margins distinctly toothed or lobed, minutely glandular, ribs weak, lobes pronounced, erect, margins ciliate. |
Capsules | included, 6–8 mm. |
included, 4–9 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
Erythranthe ptilota |
Erythranthe sierrae |
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Phenology | Flowering (May–)Jun–Aug. | Flowering Apr–Jul. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Foothill oak woodlands, mixed coniferous forests. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 200–2300 m. (700–7500 ft.) |
Distribution |
CA; OR; WA; BC
|
CA |
Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe sierrae is endemic to the southern Sierra Nevada in Fresno, Kern, and Tulare counties. The species was previously treated as E. palmeri but can be distinguished by having leaf margins that are often toothed, pale pink corollas, and white stamens. Erythranthe palmeri has entire leaf margins that are never toothed, deep pink to purple corollas, and yellow stamens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 385. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | N. S. Fraga: Aliso 30: 67, figs. 29–31. (2012) |
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