Erythranthe ptilota |
Erythranthe shevockii |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
Kelso Creek monkeyflower, Kelso Creek or Shevock's monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, taprooted. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, simple or branched from basal nodes, 2–12 cm, minutely puberulent or glabrous. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
cauline, basal not persistent; petiole 0 mm; blade palmately 3-veined (in broader ones), lanceolate to ovate, 3–10 × 1–5 mm, base truncate to truncate-cordate, clasping, margins entire, apex acute to obtuse, surfaces minutely puberulent or glabrous. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 1–16, from distal or medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
maroon and yellow, 2 lateral lobes maroon, 1 much larger central lobe yellow (red-spotted), 2 adaxial lobes maroon, bilaterally symmetric, strongly bilabiate; tube-throat cylindric, 8–12 mm, exserted 2–3 mm beyond calyx margin; limb expanded 8–15 mm, central lobe 2-fid, abaxial limb sparsely villous-bearded. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
ascending to often spreading horizontally, 10–22 mm. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
red-spotted or red, campanulate, 4–7 mm, margins distinctly toothed or lobed, minutely puberulent or glabrous, ribs weak, lobes pronounced, erect, margins glabrous. |
Capsules | included, 6–8 mm. |
included, 5–6 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
2n | = 32. |
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Erythranthe ptilota |
Erythranthe shevockii |
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Phenology | Flowering (May–)Jun–Aug. | Flowering Mar–May. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Level openings in juniper and Joshua tree woodlands. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 900–1400 m. (3000–4600 ft.) |
Distribution |
CA; OR; WA; BC
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CA
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Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe shevockii is known only from the southernmost Sierra Nevada in Kern County. It (as Mimulus shevockii) is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 389. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus shevockii |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (Heckard & Bacigalupi) N. S. Fraga: Phytoneuron 2012-39: 35. (2012) |
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