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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower

candelabrum monkey-flower, Pulsifer's monkey-flower

Habit Perennials, rhizomatous, sometimes rooting at proximal nodes. Annuals, shallowly fibrous-rooted.
Stems

prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident.

erect, straight at nodes, simple or few-branched at base, 5–12(–18) cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped.

Leaves

cauline, basal not persistent, often congested;

petiole 0 mm, rarely 1–2(–3) mm;

blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

basal and cauline;

petiole 2–9 mm, distinctly 3-veined, 2-winged;

blade palmately 3-veined, elliptic-oblong to ovate or oblanceolate, 3–14(–23) × 2–9(–15) mm, base cuneate to attenuate, margins denticulate to entire, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped.

Flowers

herkogamous, 4–10, from medial to distal nodes.

herkogamous, 1–9, from medial to distal nodes.

Styles

glabrous.

glabrous.

Corollas

yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular;

tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin;

lobe apex rounded.

yellowish, tube-throat, palate ridges, and limb yellow to pale yellow, abaxial limb red-dotted or not, bilaterally symmetric, weakly bilabiate;

tube-throat funnelform, 6–9 mm, exserted beyond calyx margin;

lobes broadly obovate-suborbicular, apex rounded.

Fruiting pedicels

(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

divergent-arcuate, 12–38 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped.

Fruiting calyces

wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate.

cylindric, ± inflated, 7–10 mm, margins distinctly toothed or lobed, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped, lobes pronounced, erect.

Capsules

included, 6–8 mm.

included, 5–8 mm.

Anthers

included, finely hirtellous to hispidulous.

included, glabrous.

2n

= 32.

Erythranthe ptilota

Erythranthe pulsiferae

Phenology Flowering (May–)Jun–Aug. Flowering Apr–Jul.
Habitat Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. Damp depressions, moist gravel, rocky flats, granite outcrops, wet meadows, lava beds, vernal pools, forest openings, commonly in or near coniferous forests, chaparral-live oak woodlands.
Elevation 0–1000(–1900) m. (0–3300(–6200) ft.) 50–1300(–2500) m. (200–4300(–8200) ft.)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
from FNA
CA; OR; WA
[WildflowerSearch map]
Discussion

Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves.

A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata.

Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom].

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe pulsiferae is characterized by minutely stipitate-glandular vestiture (lacking villous hairs), elongate internodes, persistent basal leaves, small, palmately veined, cauline leaves with short, three-veined petioles and elliptic-oblong to ovate or oblanceolate blades, divergent-arcuate pedicels, and small, all yellow, weakly bilabiate corollas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 402. FNA vol. 17, p. 398.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus moschatus var. sessilifolius Mimulus pulsiferae
Name authority G. L. Nesom: Phytoneuron 2017-17: 4. (2017) (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 39. (2012)
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