Erythranthe ptilota |
Erythranthe pulsiferae |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
candelabrum monkey-flower, Pulsifer's monkey-flower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, shallowly fibrous-rooted. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, straight at nodes, simple or few-branched at base, 5–12(–18) cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
basal and cauline; petiole 2–9 mm, distinctly 3-veined, 2-winged; blade palmately 3-veined, elliptic-oblong to ovate or oblanceolate, 3–14(–23) × 2–9(–15) mm, base cuneate to attenuate, margins denticulate to entire, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 1–9, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellowish, tube-throat, palate ridges, and limb yellow to pale yellow, abaxial limb red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 6–9 mm, exserted beyond calyx margin; lobes broadly obovate-suborbicular, apex rounded. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
divergent-arcuate, 12–38 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
cylindric, ± inflated, 7–10 mm, margins distinctly toothed or lobed, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped, lobes pronounced, erect. |
Capsules | included, 6–8 mm. |
included, 5–8 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
2n | = 32. |
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Erythranthe ptilota |
Erythranthe pulsiferae |
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Phenology | Flowering (May–)Jun–Aug. | Flowering Apr–Jul. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Damp depressions, moist gravel, rocky flats, granite outcrops, wet meadows, lava beds, vernal pools, forest openings, commonly in or near coniferous forests, chaparral-live oak woodlands. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 50–1300(–2500) m. (200–4300(–8200) ft.) |
Distribution |
CA; OR; WA; BC
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CA; OR; WA
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Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe pulsiferae is characterized by minutely stipitate-glandular vestiture (lacking villous hairs), elongate internodes, persistent basal leaves, small, palmately veined, cauline leaves with short, three-veined petioles and elliptic-oblong to ovate or oblanceolate blades, divergent-arcuate pedicels, and small, all yellow, weakly bilabiate corollas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 398. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus pulsiferae |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) |
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