Erythranthe ptilota |
Erythranthe percaulis |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
serpentine canyon monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, fibrous-rooted. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect or slightly ascending from base, often purplish, simple or few-branched from basal nodes, 7–28 cm, glabrous. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
basal and cauline; petiole: basal and proximalmost cauline 5–10 mm or cauline 0 mm; blade often spreading at right angles to stem, purple, palmately 3–5-veined, narrowly ovate, rhombic-elliptic, ovate to lanceolate, or elliptic-lanceolate to ovate or oblong-ovate, 7–10 mm, cauline even-sized or slightly smaller distally, 4–10 mm, base truncate to attenuate, margins entire or proximals shallowly sinuate, serrations 2–4, shallow, apex rounded, surfaces glabrous. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 8–12, usually on distal 2/3 of stem, not clustered, chasmogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellow, throat floor with a few red dots, proximal middle lip base with a larger red splotch, bilaterally symmetric, bilabiate; tube-throat cylindric-funnelform, 4–6 mm, exserted 2–3 mm beyond calyx margin; limb expanded 4–5 mm, palate ridges yellow, densely hairy. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
15–35 mm, glabrous. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
sharply wing-angled, urceolate to urceolate-campanulate, inflated, sagittally compressed, (4–)5–6 mm, glabrous, throat closing. |
Capsules | included, 6–8 mm. |
included, 3 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
Erythranthe ptilota |
Erythranthe percaulis |
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Phenology | Flowering (May–)Jun–Aug. | Flowering May–Jun. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Soil pockets, crevices, and boulders on serpentine cliffs, slopes, and roadcuts. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 2800 m. (9200 ft.) |
Distribution |
CA; OR; WA; BC
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CA |
Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe percaulis was described from only the type collection from Serpentine Canyon of the Feather River in Plumas County, but the type locality has recently been relocated and the population determined to comprise many thousands of individuals (S. Schoenig 2016). Plants are characterized by their completely glabrous vestiture, terete and mostly simple stems, small leaves on relatively widely spaced nodes, small calyces, and small, yellow corollas with herkogamous arrangement of stigma and anthers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 418. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | G. L. Nesom: Phytoneuron 2013-70: 1, figs. 1–5. (2013) |
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