FNA: Erythranthe ptilota vs. Erythranthe parvula

	Erythranthe ptilota	Erythranthe parvula
	musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower	Southwestern mat monkeyflower
Habit	Perennials, rhizomatous, sometimes rooting at proximal nodes.	Perennials, rhizomatous, sometimes rooting at nodes, mat-forming.
Stems	prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident.	procumbent, branched, 5–15 cm, stipitate-glandular distally.
Leaves	cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.	cauline; petiole 1–4 mm; blade palmately 3-veined, ovate to orbicular-ovate or depressed-ovate, 3–11 × 3–9 mm, base truncate to cuneate, margins shallowly denticulate to dentate, teeth 3–5 per side, apex acute to obtuse, surfaces villous-hirsute, hairs whitish, thickened, flattened, stiff, gland-tipped.
Flowers	herkogamous, 4–10, from medial to distal nodes.	plesiogamous, 2–8, axillary at distal nodes.
Styles	glabrous.	glabrous.
Corollas	yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded.	yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 6–8 mm, exserted beyond calyx margin; limb expanded 4–6 mm, abaxial limb spreading, lobes fimbriate.
Fruiting pedicels	(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.	7–15 mm, minutely stipitate-glandular.
Fruiting calyces	wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate.	nodding 45–90º, 5-lobed, cylindric-ovoid, inflated, sagittally compressed, 4–5 mm, villous-hirsute, throat closing.
Capsules	included, 6–8 mm.	included, 3–4 mm.
Anthers	included, finely hirtellous to hispidulous.	included, glabrous.
2n		= 32.

	Erythranthe ptilota	Erythranthe parvula
Phenology	Flowering (May–)Jun–Aug.	Flowering Apr–Sep.
Habitat	Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs.	Wet vertical rock faces, ledges, and rocky slopes, seepy wash banks.
Elevation	0–1000(–1900) m. (0–3300(–6200) ft.)	500–2400(–3400) m. (1600–7900(–11200) ft.)
Distribution	CA; OR; WA; BC [WildflowerSearch map]	AZ; NM; Mexico (Sonora)
Discussion	Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.)	All monkeyflowers with laciniate-lobed corollas have generally been identified as Mimulus dentilobus B. L. Robinson & Fernald, but these occur as three, morphologically distinct, widely allopatric population systems, each of which is regarded as a separate species (G. L. Nesom 2012g). Erythranthe dentiloba (B. L. Robinson & Fernald) G. L. Nesom, which is endemic to Mexico, is the only one of the three with an allogamous breeding system. Erythranthe parvula is restricted to Arizona and New Mexico except for one locality in northern Sonora, Mexico. Erythranthe chinatiensis is the third species of the group. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 402.	FNA vol. 17, p. 424.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus moschatus var. sessilifolius	Mimulus parvulus
Name authority	G. L. Nesom: Phytoneuron 2017-17: 4. (2017)	(Wooton & Standley) G. L. Nesom: Phytoneuron 2012-39: 44. (2012)
Web links	Local floras: BC, CA, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe parvula

musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower

Southwestern mat monkeyflower

Habit

Perennials, rhizomatous, sometimes rooting at proximal nodes.

Perennials, rhizomatous, sometimes rooting at nodes, mat-forming.

Stems

prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident.

procumbent, branched, 5–15 cm, stipitate-glandular distally.

Leaves

cauline, basal not persistent, often congested;

petiole 0 mm, rarely 1–2(–3) mm;

blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

cauline;

petiole 1–4 mm;

blade palmately 3-veined, ovate to orbicular-ovate or depressed-ovate, 3–11 × 3–9 mm, base truncate to cuneate, margins shallowly denticulate to dentate, teeth 3–5 per side, apex acute to obtuse, surfaces villous-hirsute, hairs whitish, thickened, flattened, stiff, gland-tipped.

Flowers

herkogamous, 4–10, from medial to distal nodes.

plesiogamous, 2–8, axillary at distal nodes.

Styles

glabrous.

Corollas

yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular;

tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin;

lobe apex rounded.

yellow, red-dotted, bilaterally symmetric, bilabiate;

tube-throat funnelform, 6–8 mm, exserted beyond calyx margin;

limb expanded 4–6 mm, abaxial limb spreading, lobes fimbriate.

Fruiting pedicels

(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

7–15 mm, minutely stipitate-glandular.

Fruiting calyces

wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate.

nodding 45–90º, 5-lobed, cylindric-ovoid, inflated, sagittally compressed, 4–5 mm, villous-hirsute, throat closing.

Capsules

included, 6–8 mm.

included, 3–4 mm.

Anthers

included, finely hirtellous to hispidulous.

included, glabrous.

= 32.

Erythranthe ptilota

Erythranthe parvula

Phenology

Flowering (May–)Jun–Aug.

Flowering Apr–Sep.

Habitat

Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs.

Wet vertical rock faces, ledges, and rocky slopes, seepy wash banks.

Elevation

0–1000(–1900) m. (0–3300(–6200) ft.)

500–2400(–3400) m. (1600–7900(–11200) ft.)

Distribution

CA; OR; WA; BC

[WildflowerSearch map]

AZ; NM; Mexico (Sonora)

Discussion

Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves.

A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata.

Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom].

(Discussion copyrighted by Flora of North America; reprinted with permission.)

All monkeyflowers with laciniate-lobed corollas have generally been identified as Mimulus dentilobus B. L. Robinson & Fernald, but these occur as three, morphologically distinct, widely allopatric population systems, each of which is regarded as a separate species (G. L. Nesom 2012g). Erythranthe dentiloba (B. L. Robinson & Fernald) G. L. Nesom, which is endemic to Mexico, is the only one of the three with an allogamous breeding system. Erythranthe parvula is restricted to Arizona and New Mexico except for one locality in northern Sonora, Mexico. Erythranthe chinatiensis is the third species of the group.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 402.

FNA vol. 17, p. 424.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus moschatus var. sessilifolius

Mimulus parvulus

Name authority

G. L. Nesom: Phytoneuron 2017-17: 4. (2017)

(Wooton & Standley) G. L. Nesom: Phytoneuron 2012-39: 44. (2012)

Web links

Local floras: BC, CA, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, UW Burke Herbarium

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Erythranthe ptilota

Erythranthe parvula

Erythranthe ptilota

Erythranthe parvula