Erythranthe ptilota |
Erythranthe montioides |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
montia-like monkeyflower, mountain monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, taprooted. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, simple or branched from basal nodes, 2–15 cm, glabrous or minutely puberulent, internodes elongate, distinct. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
cauline, basal not persistent; petiole 0 mm; blade 1-veined or palmately 3-veined (in broader ones), linear to lanceolate, (3–)4–25 × 0.5–2 mm, base attenuate, margins entire, apex acute, surfaces glabrous or minutely puberulent. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 1–6, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellow, abaxial limb red-spotted, bilaterally symmetric, strongly bilabiate; tube-throat cylindric to funnelform, 6–10 mm, exserted beyond calyx margin; limb expanded 5–12 mm, lateral lobes entire or shallowly notched, palate glabrous or sparsely bearded. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
(4–)5–20 mm. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
becoming straw colored, campanulate, 5–7 mm, margins distinctly toothed or lobed, ciliate, glabrous or minutely puberulent, ribs weak, lobes pronounced, erect. |
Capsules | included, 6–8 mm. |
included, 5–6 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
Erythranthe ptilota |
Erythranthe montioides |
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Phenology | Flowering (May–)Jun–Aug. | Flowering Jun–Aug. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Sandy opening in mixed coniferous and lodgepole forests. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 1900–2900 m. (6200–9500 ft.) |
Distribution |
CA; OR; WA; BC
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CA
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Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe montioides has been previously treated as a highly polymorphic species with a relatively broad geographic range but is now recognized as narrowly endemic to Fresno and north-central Tulare counties. The following species were segregated from E. montioides: E. barbata, E. calcicola, E. carsonensis, and E. discolor. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 385. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus montioides |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (A. Gray) N. S. Fraga: Phytoneuron 2012-39: 35. (2012) |
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