Erythranthe ptilota |
Erythranthe minor |
|
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
Colorado monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Perennials, rhizomatous, colonial, rhizomes forming a mass, branching, filiform. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect to erect-ascending, branched, 5–20 cm, densely minutely hirtellous and eglandular or with a mixture of hirtellous and gland-tipped hairs. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
basal and cauline; petiole 0 mm or proximals 1–3 mm; blade palmately 3-veined, broadly ovate to elliptic-ovate or lanceolate, 8–25 × 5–15 mm, base cuneate to truncate, margins shallowly dentate to denticulate, apex acute to obtuse, surfaces glabrous. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 1–3, from distal nodes. |
Styles | glabrous. |
sparsely hirtellous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellow, not red-dotted, bilaterally symmetric, bilabiate; tube-throat tubular-funnelform, 9–11 mm, exserted 0–1(–2) mm beyond calyx margin. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
10–20 mm, densely minutely hirtellous and eglandular or with a mixture of hirtellous and gland-tipped hairs. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
nodding 80–100º, not purple-dotted, cylindric-campanulate, inflated, sagittally compressed, 10–13 mm, densely minutely hirtellous and eglandular or with a mixture of hirtellous and gland-tipped hairs, throat closing. |
Capsules | included, 6–8 mm. |
included, 5–8 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
Erythranthe ptilota |
Erythranthe minor |
|
Phenology | Flowering (May–)Jun–Aug. | Flowering Jul–Aug(–Sep). |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Stream and lake edges, intermittent subalpine water courses, roadside ditches, subalpine to alpine. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 3000–3700 m. (9800–12100 ft.) |
Distribution |
CA; OR; WA; BC
|
CO; NM |
Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The corollas of Erythranthe minor are shorter than those of typical E. tilingii, and the two species are allopatric. Corollas of E. tilingii rarely may be equally as short as those of E. minor but are produced in scattered localities on plants that are depauperate in other ways. The range of E. minor is primarily in Colorado apparently extending southward into the Wheeler Peak area of Taos County, New Mexico. Attribution of its range into the La Sal Mountains of east-central Utah has been based on misidentifications of E. guttata; the distinction between E. guttata and E. minor in Colorado also needs clarification. Mimulus luteus Linnaeus var. alpinus A. Gray (1863, the type from Colorado) is an illegitimate name for Erythranthe minor, preceded by M. luteus var. alpinus Lindley (1827). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 409. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus minor, M. alpinus, M. langsdorffii var. alpinus, M. langsdorffii var. minor |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (A. Nelson) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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