Erythranthe ptilota |
Erythranthe linearifolia |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
primrose monkeyflower, threadleaf primrose monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Perennials, rhizomatous, densely cespitose, forming large patches and turfs 0.3–1 m diam. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect to ascending, simple, 2–10 cm, sparsely hirsute and stipitate-glandular, internodes shortened. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
basal or near basal, sometimes proximal cauline, subrosulate; petiole 0 mm; blade 1-veined or palmately 3-veined, linear to narrowly oblanceolate, 15–50 × 1.5–5 mm, base long-cuneate, often subclasping, margins entire, dentate-serrate, or distally dentate, apex acute, surfaces glabrous or adaxial sparsely short-pilose, eglandular. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 1. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellow, red-spotted or -striped, bilaterally symmetric, weakly bilabiate, loosely hirsute on abaxial side of opening; tube-throat narrowly campanulate, 18–22 mm, exserted beyond calyx margin; lobes broadly obovate-oblong, apex rounded- or truncate-notched, throat open. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
(40–)65–85(–120) mm, glabrous or sparsely stipitate-glandular near base. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
winged- or plicate-angled, tubular-campanulate, weakly or not inflated, 9–10(–12) mm, glabrous. |
Capsules | included, 6–8 mm. |
included, 6–7 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included or slightly exserted, margins ciliate, glabrous. |
Erythranthe ptilota |
Erythranthe linearifolia |
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Phenology | Flowering (May–)Jun–Aug. | Flowering Jul–Sep. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Wet banks, Darlingtonia seeps and bogs, seepages in serpentine talus. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 600–2800 m. (2000–9200 ft.) |
Distribution |
CA; OR; WA; BC
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CA
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Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe linearifolia is endemic to serpentine substrates in Shasta, Siskiyou, and Trinity counties; typical E. primuloides occurs in the same area but not on serpentine. Erythranthe linearifolia is distinct from E. primuloides especially in its narrow leaves and cespitose habit. A collection from Tulare County appears to be E. linearifolia (Shevock 10597, CAS), but this appears to be far out of range and the voucher should be reexamined; it probably is better identified as an unusual collection of E. primuloides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 390. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus primuloides var. linearifolius, M. linearifolius, M. primuloides subsp. linearifolius |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (A. L. Grant) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 35. (2012) |
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