Erythranthe ptilota |
Erythranthe lewisii |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
great purple monkey-flower, great purple or Lewis' monkeyflower, Lewis' monkey flower, pink monkey-flower, purple monkey-flower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Perennials, rhizomatous. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, usually simple, (15–)25–60(–75) cm, stipitate-glandular to glandular-villous. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
cauline; petiole 0 mm; blade palmately veined, elliptic to ovate, ovate-lanceolate, or broadly oblanceolate, (10–)25–75(–90) × 5–35 mm, base rounded to cuneate, subclasping, margins denticulate, subentire, or entire, apex acute, surfaces stipitate-glandular to glandular-villous. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 2–6(–10), axillary at leafy medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
purple, rarely crimson, pale violet, white, pinkish white, yellowish white, or lavender, sometimes lined with red dots, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 22–28 mm, exserted beyond calyx margins; lobe apex usually truncate to shallowly convex, shallowly retuse, throat open. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
(25–)35–70 mm, stipitate-glandular to glandular-villous. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
broadly cylindric-campanulate, not inflated, 15–22 mm, stipitate-glandular to glandular-villous, tube 12–15(–17) × 9–12 mm. |
Capsules | included, 6–8 mm. |
included, 6–11 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, white-villous, thecae spreading. |
2n | = 16. |
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Erythranthe ptilota |
Erythranthe lewisii |
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Phenology | Flowering (May–)Jun–Aug. | Flowering Jun–Sep. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Stream banks, springs, wet meadows, subalpine slopes, talus, crevices, ditches. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 600–2900(–3200) m. (2000–9500(–10500) ft.) |
Distribution |
CA; OR; WA; BC
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AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
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Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe lewisii in California occurs in Modoc, Siskiyou, and Trinity counties. California records from Shasta County and south previously identified as E. lewisii are identified here as E. erubescens. The record for E. lewisii in Alaska is documented by this collection: Hyder [noted on handwritten label as "New to Alaska, Standley"], damp land, 27 June 1924, K. Whited 1291 (MO). Apparent exceptions to the characteristic flower color are these: white to lavender in Nevada (Clark County, Charleston Mountains, Train 2068, MO); pinkish white in Washington (Skamania and Yakima counties, Mt. Paddo, Suksdorf 5779, MO); white or tinged with yellow, in Wyoming (Teton County, as described by Nelson in the protologue of Mimulus lewisii var. tetonensis). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 392. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus lewisii, M. lewisii var. tetonensis |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (Pursh) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) |
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