Erythranthe ptilota |
Erythranthe hallii |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
Hall's monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, fibrous-rooted, sometimes apparently rooting at proximal nodes if stems proximally decumbent. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, simple, 4-angled, 2–8 cm, slender, glabrous. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
basal and cauline or basal deciduous, largest at mid stem or above, cauline relatively few on long internodes; petiole: basal and proximals to midcauline 1–4 mm, distals 0 mm; blade palmately 3-veined, ovate to ovate-lanceolate, 5–11 × 3–9 mm, base truncate to cuneate, margins very shallowly dentate or denticulate, apex acute to obtuse, surfaces glabrous or distals and bracteals sparsely villous, hairs vitreous, flattened, eglandular, multicellular. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
plesiogamous, (1–)4–10, sometimes from all nodes, usually beginning about mid stem, cleistogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellow, usually red-dotted, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly cylindric, 4–6 mm, exserted 0.5–1 mm beyond calyx margin; limb barely expanded. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
usually deflexed 90º at calyx, 6–14 mm, longer than subtending leaves. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
sometimes red-dotted, broadly elliptic-ovoid, inflated, sagittally compressed, (5–)7–10 mm, glabrous, throat closing, adaxial lobe not distinctly longer than abaxial, not falcate. |
Capsules | included, 6–8 mm. |
included, 4–6 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
2n | = 32. |
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Erythranthe ptilota |
Erythranthe hallii |
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Phenology | Flowering (May–)Jun–Aug. | Flowering May–Aug. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Ledges, seeps, along streams, wet meadows. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 1900–3200 m. (6200–10500 ft.) |
Distribution |
CA; OR; WA; BC
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CO |
Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe hallii is known from Boulder, Clear Creek, Fremont, Grand, Jefferson, Larimer, Routt, and Saguache counties. The Colorado population system is morphologically and geographically coherent. Erythranthe hallii is similar to E. arvensis; both have four-angled stems, autogamous reproduction, a tendency to root at basal nodes and distally, and both have bracteal leaves villous with vitreous, flattened, eglandular, multicellular hairs, although this vestiture is barely developed and often absent in E. hallii. The only reported chromosome number from the Colorado plants (2n = 32) also appears to be distinct among possible relatives of E. hallii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 423. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus hallii, M. guttatus var. hallii |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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