Erythranthe ptilota |
Erythranthe filicifolia |
|
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
fern-leaf monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, fibrous-rooted. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect or slightly ascending from base, sometimes purplish, simple or few-branched from proximal nodes, 3–38 cm, glabrous. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
basal and cauline; petiole (proximal and proximal to mid cauline), distal bracteate or absent, ovate, margins entire; blade: divisions 1-veined, oblong-lanceolate to ovate in outline, 3–68 mm, slightly thickened, base (at proximalmost division) truncate, margins (of divisions) entire, bipinnately dissected, ca. 5–12(–15) primary divisions on each side, ultimate division nearly linear, apex obtuse to acute, surfaces glabrous. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
plesiogamous, 4–10(–12), on foreshortened distal nodes, usually clustered, chasmogamous, sometimes cleistogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellow, proximal lip sometimes red-splotched, palate ridges yellow, bilaterally symmetric, bilabiate; tube-throat cylindric-funnelform, 4–8 mm, exserted 0–2 mm beyond calyx margin or not; palate ridges densely villous. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
6–14 mm, glabrous. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
urceolate to urceolate-campanulate, inflated, sagittally compressed, 8–11 mm, glabrous, throat closing. |
Capsules | included, 6–8 mm. |
included, 3–8 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
Erythranthe ptilota |
Erythranthe filicifolia |
|
Phenology | Flowering (May–)Jun–Aug. | Flowering Apr–Jun(–Sep). |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Slow-draining, ephemeral seeps on exfoliating granite slabs, over basalt. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 300–1600 m. (1000–5200 ft.) |
Distribution |
CA; OR; WA; BC
|
CA |
Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe filicifolia is known only from localities in Butte and Plumas counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 418. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus filicifolius |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (Sexton: K. G. Ferris & Schoenig) G. L. Nesom, Phytoneuron 2013-80: 1. (2013) |
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