Erythranthe ptilota |
Erythranthe filicaulis |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
Hetch Hetchy monkey-flower, slender-stem monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, taprooted. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, simple or branched at basal nodes, 4–30 cm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
cauline, basal usually not persistent; petiole 0 mm; blade 1-veined or palmately 3-veined, linear to oblanceolate, 6–15 × (1–)2–3 mm, base attenuate, margins entire to remotely mucronulate, teeth 1–3 per side, apex acute, surfaces densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 2–8, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
pink to purple, throat dark red-purple, palate ridges yellow, bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat cylindric-funnelform, 8–12(–17) mm, exserted beyond calyx margin; limb expanded 10–14 mm, lobes notched, palate villous. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
9–30 mm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
reddish brown-spotted, strongly angled, cylindric-campanulate, 7–9 mm, margins distinctly toothed or lobed, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped, ribs corky, lobes pronounced, spreading. |
Capsules | included, 6–8 mm. |
included, 5–7 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, hirtellous. |
2n | = 16. |
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Erythranthe ptilota |
Erythranthe filicaulis |
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Phenology | Flowering (May–)Jun–Aug. | Flowering May–Jul. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Moist open areas on gentle slopes, meadows, roadsides, gravelly soils. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 800–1800 m. (2600–5900 ft.) |
Distribution |
CA; OR; WA; BC
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CA
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Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe filicaulis is known only for certain from Mariposa and Tuolumne counties. Kappler 1691 (UCLA) from Madera County also may be this species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 391. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus filicaulis, M. biolettii |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (S. Watson) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) |
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