Erythranthe ptilota |
Erythranthe eastwoodiae |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
crimson monkeyflower, Eastwood's monkey-flower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Perennials, stoloniferous, sometimes also rhizomatous. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
scandent to pendent, usually simple, 5–30(–40) cm, villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
cauline; petiole 0 mm; blade palmately 3-veined, flabellate distally to obovate to oblanceolate or elliptic, (5–)13–40(–55) × 8–20(–25) mm, largest near mid stem or distally, thick, base cuneate to rounded, subclasping, margins coarsely serrate on distal 1/2, apex acute, surfaces villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 2–8, axillary at leafy medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
scarlet to orange-red or orange, palate red, not spotted or striped, bilaterally symmetric, strongly bilabiate; tube-throat narrowly funnelform, 20–30 mm, exserted 5–15 mm beyond calyx margin; throat open, palate puberulent. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
10–30(–40) mm. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
cuneate-cylindric to cylindric, weakly or not inflated, 15–23(–27) mm, glabrous or minutely stipitate-glandular to sparsely glandular-villosulous, lobes triangular-acuminate. |
Capsules | included, 6–8 mm. |
included, 6–10 mm. |
Anthers | included, finely hirtellous to hispidulous. |
exserted, villous, thecae reflexed 45º. |
2n | = 16. |
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Erythranthe ptilota |
Erythranthe eastwoodiae |
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Phenology | Flowering (May–)Jun–Aug. | Flowering May–Sep(–Nov). |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Seepages in sandstone overhangs, cave roofs, walls, crevices, and cliff bases, pinyon-juniper woodlands. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 900–2000 m. (3000–6600 ft.) |
Distribution |
CA; OR; WA; BC
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AZ; CO; NM; UT
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Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The range of Erythranthe eastwoodiae appears to be essentially contiguous with that of E. verbenacea in the Grand Canyon region, but there is no evidence of hybridization. Erythranthe eastwoodiae (as Mimulus eastwoodiae) is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 394. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus eastwoodiae |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (Rydberg) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) |
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