FNA: Erythranthe ptilota vs. Erythranthe chinatiensis

	Erythranthe ptilota	Erythranthe chinatiensis
	musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower	Chinati Mountains monkeyflower
Habit	Perennials, rhizomatous, sometimes rooting at proximal nodes.	Perennials, rhizomatous, sometimes rooting at nodes, mat-forming.
Stems	prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident.	procumbent, branched, 5–20 cm, glabrous.
Leaves	cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.	cauline; petiole 2–10(–20) mm; blade palmately 3–5(–7)-veined, ovate to broadly ovate or orbicular-ovate, 4–15(–22) × 4–15(–18) mm, base truncate to cuneate, margins shallowly denticulate or merely mucronate to mucronulate, teeth 3–6 per side, apex acute to obtuse, surfaces glabrous, adaxial sometimes moderately villosulous, hairs vitreous, flattened, eglandular or minutely gland-tipped.
Flowers	herkogamous, 4–10, from medial to distal nodes.	plesiogamous, 2–8, axillary at distal nodes.
Styles	glabrous.	glabrous.
Corollas	yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded.	yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 7–8 mm, exserted beyond calyx margin; limb expanded 6–7 mm, abaxial limb strongly reflexed, lobes fimbriate.
Fruiting pedicels	(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.	10–20 mm, glabrous.
Fruiting calyces	wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate.	nodding 45–90º, 5-lobed, ellipsoid, inflated, sagittally compressed, 5–6 mm, glabrous or sparsely villosulous-glandular, throat closing.
Capsules	included, 6–8 mm.	included, 4–5 mm.
Anthers	included, finely hirtellous to hispidulous.	included, glabrous.

	Erythranthe ptilota	Erythranthe chinatiensis
Phenology	Flowering (May–)Jun–Aug.	Flowering Feb–Sep.
Habitat	Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs.	Seeps in vertical cliff faces, wet bluffs.
Elevation	0–1000(–1900) m. (0–3300(–6200) ft.)	600–1900(–2300) m. (2000–6200(–7500) ft.)
Distribution	CA; OR; WA; BC [WildflowerSearch map]	TX
Discussion	Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe chinatiensis is similar to E. parvula in its prostrate habit, five-lobed calyces, and fimbriate corolla lobes. It differs from the latter in its nearly glabrous leaves and strongly reflexed abaxial corolla lip. Erythranthe chinatiensis is known only from Presidio County but should be expected to occur also in adjacent Chihuahua, Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 402.	FNA vol. 17, p. 424.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus moschatus var. sessilifolius
Name authority	G. L. Nesom: Phytoneuron 2017-17: 4. (2017)	G. L. Nesom: Phytoneuron 2012-40: 86, figs. 12–14. (2012)
Web links	Local floras: BC, CA, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe chinatiensis

musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower

Chinati Mountains monkeyflower

Habit

Perennials, rhizomatous, sometimes rooting at proximal nodes.

Perennials, rhizomatous, sometimes rooting at nodes, mat-forming.

Stems

prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident.

procumbent, branched, 5–20 cm, glabrous.

Leaves

cauline, basal not persistent, often congested;

petiole 0 mm, rarely 1–2(–3) mm;

blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

cauline;

petiole 2–10(–20) mm;

blade palmately 3–5(–7)-veined, ovate to broadly ovate or orbicular-ovate, 4–15(–22) × 4–15(–18) mm, base truncate to cuneate, margins shallowly denticulate or merely mucronate to mucronulate, teeth 3–6 per side, apex acute to obtuse, surfaces glabrous, adaxial sometimes moderately villosulous, hairs vitreous, flattened, eglandular or minutely gland-tipped.

Flowers

herkogamous, 4–10, from medial to distal nodes.

plesiogamous, 2–8, axillary at distal nodes.

Styles

glabrous.

Corollas

yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular;

tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin;

lobe apex rounded.

yellow, red-dotted, bilaterally symmetric, bilabiate;

tube-throat funnelform, 7–8 mm, exserted beyond calyx margin;

limb expanded 6–7 mm, abaxial limb strongly reflexed, lobes fimbriate.

Fruiting pedicels

(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

10–20 mm, glabrous.

Fruiting calyces

wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate.

nodding 45–90º, 5-lobed, ellipsoid, inflated, sagittally compressed, 5–6 mm, glabrous or sparsely villosulous-glandular, throat closing.

Capsules

included, 6–8 mm.

included, 4–5 mm.

Anthers

included, finely hirtellous to hispidulous.

included, glabrous.

Erythranthe ptilota

Erythranthe chinatiensis

Phenology

Flowering (May–)Jun–Aug.

Flowering Feb–Sep.

Habitat

Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs.

Seeps in vertical cliff faces, wet bluffs.

Elevation

0–1000(–1900) m. (0–3300(–6200) ft.)

600–1900(–2300) m. (2000–6200(–7500) ft.)

Distribution

CA; OR; WA; BC

[WildflowerSearch map]

Discussion

Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves.

A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata.

Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom].

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe chinatiensis is similar to E. parvula in its prostrate habit, five-lobed calyces, and fimbriate corolla lobes. It differs from the latter in its nearly glabrous leaves and strongly reflexed abaxial corolla lip. Erythranthe chinatiensis is known only from Presidio County but should be expected to occur also in adjacent Chihuahua, Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 402.

FNA vol. 17, p. 424.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus moschatus var. sessilifolius

Name authority

G. L. Nesom: Phytoneuron 2017-17: 4. (2017)

G. L. Nesom: Phytoneuron 2012-40: 86, figs. 12–14. (2012)

Web links

Local floras: BC, CA, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, UW Burke Herbarium

The species comparison tool is brought to you by:

Flora of North America species comparison

Erythranthe ptilota

Erythranthe chinatiensis

Erythranthe ptilota

Erythranthe chinatiensis