Erythranthe ptilota |
Erythranthe carsonensis |
|
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
Carson Valley monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, taprooted, densely compact. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, simple or branched from basal nodes, 2–7(–8) cm, minutely glandular, internodes shortened, not evident. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
cauline, basal not persistent; petiole 0 mm; blade palmately 3-veined (in broader ones), linear to spatulate, (3–)5–23 × 1–5 mm, base truncate to truncate-cordate, clasping, margins entire, apex acute to obtuse, surfaces minutely glandular. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 1–35, from distal or medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellow, palate red-dotted and 1 large central spot, bilaterally symmetric, strongly bilabiate; tube-throat cylindric, distinct from abruptly expanding throat, (5–)7–11 mm, exserted beyond calyx margin; limb expanded 7–12(–15) mm, each lobe 2-fid, palate densely bearded. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
(3–)5–14 mm. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
campanulate to widely urceolate, 4–7 mm, margins distinctly toothed or lobed, minutely glandular, ribs thickened, lobes pronounced, erect. |
Capsules | included, 6–8 mm. |
included, 3–6 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
Erythranthe ptilota |
Erythranthe carsonensis |
|
Phenology | Flowering (May–)Jun–Aug. | Flowering Mar–Jun. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Openings in sage brush/bitterbrush scrub in sand of decomposed granite. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 1400–1800 m. (4600–5900 ft.) |
Distribution |
CA; OR; WA; BC
|
CA; NV |
Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe carsonensis is restricted to the Carson Valley, Eagle Valley, and Washoe Valley region of Nevada and adjacent California, with one known disjunct occurrence about 58 km to the north in Nevada. The species was previously included in the broader concept of E. montioides but can be distinguished by its much branched and compact habit, linear to spatulate leaves with clasping bases, calyx with glabrous margins, and larger corolla with one large red spot in the center. Erythranthe carsonensis has been impacted by agriculture, urbanization, and other anthropogenic changes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 384. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus rubellus var. latiflorus |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | N. S. Fraga: Aliso 30: 59, figs. 17–21. (2012) |
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