Erythranthe ptilota |
Erythranthe calciphila |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
Mexican moss monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, usually fibrous-rooted, rarely rooting at proximal nodes. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, sometimes decumbent-ascending, branched, 4–30 cm, minutely stipitate-glandular, also delicately villosulous-glandular along whole length. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
basal and cauline; petiole 1–3 mm, cauline blade slightly or hardly reduced in size from basal, becoming subsessile to sessile (1–3 pairs of cauline leaves); blade palmately 3(–5)-veined, orbicular-ovate to oblong-ovate, 7–28 × 5–22 mm, base truncate to shallowly cuneate, margins shallowly dentate to denticulate, teeth 3–6 per side, apex rounded to obtuse, surfaces villous, hairs thin-walled, vitreous, eglandular or minutely gland-tipped, usually also minutely stipitate-glandular. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
plesiogamous, 1–6(–10), axillary at all nodes, chasmogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
light yellow, red-spotted, weakly bilaterally symmetric, ± bilabiate; tube-throat narrowly funnelform, 5–7 mm, exserted 1–2 mm beyond calyx margin; limb expanded 2–4 mm. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
15–30(–55) mm, minutely stipitate-glandular, sometimes minutely hirtellous and minutely stipitate-glandular, sometimes short glandular-villous. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
nodding 90º at maturity, usually 3(–5)-veined, ovoid, inflated, sagittally compressed, 6–10 mm, minutely stipitate-glandular, throat closing, lobes usually 3 or 3 and 5 on same plant, if 5 then with 2 interpolated lobes much smaller than abaxial pair. |
Capsules | included, 6–8 mm. |
included, 2–4 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
2n | = 30, 32. |
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Erythranthe ptilota |
Erythranthe calciphila |
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Phenology | Flowering (May–)Jun–Aug. | Flowering Mar–Sep(–Nov). |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Rocky knobs, moist boulders, wet rock faces, roadcuts, seepages, springs, with moss, usually in pine or pine-oak woods. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 1800–2500 m. (5900–8200 ft.) |
Distribution |
CA; OR; WA; BC
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AZ; Mexico (Chihuahua, Durango, Sinaloa, Sonora) |
Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe calciphila is recognized by its annual duration (fibrous-rooted), short, erect stems with few, even-sized leaves (the basal often persistent), delicate stipitate-glandular vestiture, three-lobed calyces that are relatively large in fruit, small corollas, and autogamous reproduction. Plants rarely root at proximal nodes. In Arizona, at the northernmost extremity of its range, E. calciphila occurs in the Chiricahua, Huachuca, and Mule mountains of Cochise County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 424. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus calciphilus, M. minutiflorus |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (Gentry) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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