Erythranthe ptilota |
Erythranthe calcicola |
|
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
limestone monkeyflower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, taprooted. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, simple or branched from basal nodes, 2–10(–15) cm, sparsely glandular-pubescent, internodes elongate, distinct. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
cauline, basal not persistent; petiole 0–1 mm; blade palmately 3-veined (in broader ones), lanceolate to ovate, 3–25 × 2–8(–10) mm, base attenuate, margins entire, sometimes toothed, apex acute to obtuse, surfaces sparsely glandular-pubescent. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, sometimes plesiogamous, 1–16, from distal or medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellow or white and throat yellow, throat and limb red-spotted, bilaterally symmetric, weakly bilabiate; tube-throat cylindric, 6–13 mm, exserted beyond calyx margin; limb expanded 3–7(–9) mm, lobes notched, abaxial limb sparsely bearded or glabrous. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
(3–)5–20 mm. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
campanulate to widely cylindric, 5–7 mm, margins distinctly toothed or lobed, sparsely glandular-pubescent, ribs strongly angled, lobes pronounced, erect. |
Capsules | included, 6–8 mm. |
included, 4–6 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
Erythranthe ptilota |
Erythranthe calcicola |
|
Phenology | Flowering (May–)Jun–Aug. | Flowering Apr–Jun. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Creosote bush and Joshua tree woodlands, usually on carbonate substrate, primarily on talus slopes. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 900–2200 m. (3000–7200 ft.) |
Distribution |
CA; OR; WA; BC
|
CA; NV |
Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe calcicola was previously included in the broader concept of E. montioides; the former can be distinguished by its glandular herbage, broader lanceolate to ovate leaves, strongly angled calyces, and smaller flowers. The species is restricted to the northern Mojave Desert and southwestern Great Basin and occurs primarily on talus slopes on substrates derived from carbonate rock. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 383. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | N. S. Fraga & D. A. York: Aliso 30: 54, figs. 12–16. (2012) |
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