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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower

bearded monkeyflower

Habit Perennials, rhizomatous, sometimes rooting at proximal nodes. Annuals, taprooted.
Stems

prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident.

erect, simple or branched from basal nodes, 2–14.5 cm, sparsely glandular-pubescent, internodes elongate, distinct.

Leaves

cauline, basal not persistent, often congested;

petiole 0 mm, rarely 1–2(–3) mm;

blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

cauline, basal not persistent;

petiole 0 mm;

blade 1-veined or palmately 3-veined (in broader ones), linear to lanceolate, 5–15 × 0.5–2 mm, base attenuate, margins entire, apex acute to obtuse, surfaces sparsely glandular-pubescent.

Flowers

herkogamous, 4–10, from medial to distal nodes.

herkogamous, 1–30, from distal or medial to distal nodes.

Styles

glabrous.

glabrous.

Corollas

yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular;

tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin;

lobe apex rounded.

bicolored (abaxial limb yellow with red spots, adaxial maroon-purple) or yellow, bilaterally symmetric, strongly bilabiate;

tube-throat cylindric, (5–)8–15 mm, exserted beyond calyx margin;

limb expanded 6–15 mm, lateral lobes 2-fid, palate bearded.

Fruiting pedicels

(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones.

ascending to often spreading horizontally, (5–)9–25 mm.

Fruiting calyces

wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate.

sometimes red-dotted on ribs, campanulate, 3–6 mm, margins distinctly toothed or lobed, glabrous, sparsely glandular-pubescent, ribs weak, lobes pronounced, erect, margins glabrous.

Capsules

included, 6–8 mm.

included or equal to calyx, 3–5 mm.

Anthers

included, finely hirtellous to hispidulous.

included, glabrous.

Erythranthe ptilota

Erythranthe barbata

Phenology Flowering (May–)Jun–Aug. Flowering May–Aug.
Habitat Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. Open areas in pine forests, edges of meadows and ephemeral streams.
Elevation 0–1000(–1900) m. (0–3300(–6200) ft.) 1800–3400 m. (5900–11200 ft.)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
from FNA
CA
[WildflowerSearch map]
Discussion

Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves.

A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata.

Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom].

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe barbata previously has been placed in synonymy with E. montioides but differs from it in having each corolla lobe deeply notched and a consistently bearded palate. In E. montioides, each corolla lobe is entire or shallowly notched, and the palate is glabrous or sparsely bearded. Erythranthe barbata is most abundant in Tulare County but also occurs in immediately adjacent Inyo and Kern counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 402. FNA vol. 17, p. 386.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus moschatus var. sessilifolius Mimulus barbatus, M. deflexus
Name authority G. L. Nesom: Phytoneuron 2017-17: 4. (2017) (Greene) N. S. Fraga: Phytoneuron 2012-39: 35. (2012)
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