Erythranthe ptilota |
Erythranthe alsinoides |
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musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
chickweed monkey-flower, wing-stem monkey-flower |
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Habit | Perennials, rhizomatous, sometimes rooting at proximal nodes. | Annuals, fibrous-rooted. |
Stems | prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
erect, usually simple, (0.5–)2–6(–15) cm, glandular-puberulent, hairs 0.1–0.2 mm, gland-tipped, nodes 2(or 3), usually red-tinged. |
Leaves | cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
basal and cauline; petiole 1–20(–30) mm, distinctly 3-veined (winged); blade palmately 3-veined, lanceolate-ovate to ovate, elliptic, or suborbicular, 3–18(–32) × 3–12(–25) mm, base cuneate to truncate, margins dentate to denticulate or subentire, apex acute to obtuse or rounded, surfaces glandular-puberulent, hairs 0.1–0.2 mm, gland-tipped. |
Flowers | herkogamous, 4–10, from medial to distal nodes. |
herkogamous, 1–4(–8), from distal or medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
yellow, abaxial limb with a large maroon splotch, also red-spotted, bilaterally symmetric, ± bilabiate; tube-throat funnelform-cylindric, 6–9 mm, exserted beyond calyx margin; throat open, palate villous, abaxial ridges low. |
Fruiting pedicels | (15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
15–32 mm, glandular-puberulent, hairs 0.1–0.2 mm, gland-tipped. |
Fruiting calyces | wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
purplish, slightly ridge-angled, campanulate-cylindric, weakly or not inflated, 5–8 mm, margins subtruncate, sparsely minutely stipitate-glandular, lobes 4, (0–)0.5–1 mm, sometimes barely evident, 1 lobe usually slightly longer, margins appearing subtruncate, shallowly convex to rounded-mucronulate. |
Capsules | included, 6–8 mm. |
included, (3–)5–7 mm. |
Anthers | included, finely hirtellous to hispidulous. |
included, glabrous. |
Erythranthe ptilota |
Erythranthe alsinoides |
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Phenology | Flowering (May–)Jun–Aug. | Flowering Mar–Jun. |
Habitat | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. | Open, rocky slopes, cliff faces, bluffs, mossy rock crevices, ledges, moist rocks, roadsides, along wet paths and trails. |
Elevation | 0–1000(–1900) m. (0–3300(–6200) ft.) | 10–1900 m. (0–6200 ft.) |
Distribution |
CA; OR; WA; BC
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CA; ID; OR; WA; BC
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Discussion | Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe alsinoides is distinct in its short, erect stems with few nodes, small, mostly ovate to elliptic-ovate, petiolate leaves, minutely stipitate-glandular vestiture, small corollas with a prominent maroon splotch on the abaxial limb, small, non-inflated mature calyces and, most especially, by its nearly truncate calyx margin. Erythranthe pulsiferae is superficially similar to E. alsinoides but has larger calyces borne on divergent-arcuate pedicels, smaller leaf blades with attenuate to cuneate bases, and the corolla limbs are smaller and lack a prominent maroon splotch. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 402. | FNA vol. 17, p. 395. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus var. sessilifolius | Mimulus alsinoides |
Name authority | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) | (Douglas ex Bentham) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 37. (2012) |
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