Erythranthe primuloides |
Erythranthe scouleri |
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primrose monkey-flower |
Columbia River monkey-flower, Scouler's monkey-flower |
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Habit | Perennials, rhizomatous or stoloniferous, mat-forming, rhizomes or stolons flagelliform. | Perennials, rhizomatous, producing long, sparsely leafy runners from basal nodes. |
Stems | erect to ascending, usually simple, 2–10(–20) cm, villous, internodes shortened. |
erect, simple or few-branched, 15–80 cm, glabrous. |
Leaves | all basal or near basal, often rosulate; petiole 0 mm; blade palmately 3-veined, oblanceolate to elliptic-obovate, 7–40 × 4–12 mm, base cuneate to attenuate, margins entire, distally denticulate to dentate, or sharply serrate-dentate, apex acute to obtuse, abaxial surface glabrous, adaxial glabrous or glabrate to sparsely to densely long-villous, eglandular. |
cauline, basal usually not persistent; petiole: proximals and medial 10–25 mm (gradually merging into blade), distals 0 mm; blade palmately (3–)5–7-veined to subpinnate, (proximal to medial) oblong-elliptic to oblong-lanceolate, 25–60 × 8–18 mm, usually 3–4 times longer than wide, base attenuate, margins evenly, shallowly dentate or crenate to mucronate or mucronulate, teeth 10–20 per side, sometimes more deeply toothed at base, apex obtuse to acute, surfaces glabrous. |
Flowers | herkogamous, 1. |
herkogamous, (1 or)2–8, from distal nodes. |
Styles | glabrous. |
minutely, prominently hirsutulous to villosulous. |
Corollas | yellow to orange-yellow, usually brown-spotted abaxially, base of each abaxial lobe usually with a larger reddish brown spot, bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular, densely hirsute on abaxial side of opening; tube-throat narrowly campanulate, 15–20 mm, exserted beyond calyx margin; lobes broadly obovate-oblong, apex rounded- or truncate-notched, throat open, palate densely villous, abaxial ridges prominent. |
yellow, without red markings, bilaterally symmetric, bilabiate; tube-throat funnelform, 20–24 mm, exserted 10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
Fruiting pedicels | 30–110(–130) mm, glabrous or sparsely stipitate-glandular near base. |
20–25 mm, glabrous. |
Fruiting calyces | tubular-campanulate, weakly or not inflated, 6–8 mm, glabrous. |
ovoid, inflated, sagittally compressed, 13–14 mm, glabrous, throat closing. |
Capsules | included, 6–7 mm. |
unknown. |
Anthers | slightly exserted, margins ciliate, glabrous. |
included, glabrous. |
2n | = 34. |
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Erythranthe primuloides |
Erythranthe scouleri |
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Phenology | Flowering Jun–Aug. | Flowering May–Aug. |
Habitat | Wet meadows, seeps, streamsides. | Salt marshes, river banks. |
Elevation | 600–3400 m. (2000–11200 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
AZ; CA; ID; MT; NM; NV; OR; UT; WA
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OR; WA |
Discussion | Flowers in Erythranthe primuloides and E. linearifolia characteristically appear to be scapose, but the scapes are pedicels arising from axils of greatly foreshortened stems. Occasionally in both species the internodes may lengthen somewhat, and the leaves are not so densely clustered at the base of the stems. In northern Klamath, western Deschutes, and eastern Douglas counties, Oregon, an area within the range of typical populations, Erythranthe primuloides has distinctively large corollas (limbs 10–15 mm wide). Apparent clones of large-flowered and smaller-flowered plants sometimes grow in close proximity or even intermixed, appearing as two different entities. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe scouleri is distinctive in its oblong-elliptic leaves with long-tapering bases and closely toothed margins, completely glabrous vestiture, long, leafy runners from basal cauline nodes, large corollas with broad limbs, and prominently hairy styles. Several features suggest a close relationship to E. decora, particularly its very large corollas, hairy styles, closely toothed leaf margins, tall, simple, and erect stems, numerous runners, and its geographic range. All collections apparently have been made near the mouth of the Columbia River in Clatsop and Columbia counties, Oregon (G. L. Nesom 2013d). Recent observations (Alexander John Wright, pers. comm.; photos http://www.inaturalist.org/observations/3537008) indicate that it also occurs in Wahkiakum County, Washington, in the Julia Butler Hansen Refuge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 389. | FNA vol. 17, p. 415. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus primuloides, M. nevadensis, M. pilosellus, M. primuloides var. minimus, M. primuloides var. pilosellus | Mimulus scouleri, M. guttatus subsp. scouleri |
Name authority | (Bentham) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 35. (2012) | (Hooker) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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