Erythranthe primuloides |
Erythranthe marmorata |
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primrose monkey-flower |
Stanislaus monkeyflower, Whipple's monkeyflower |
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Habit | Perennials, rhizomatous or stoloniferous, mat-forming, rhizomes or stolons flagelliform. | Annuals, taprooted. |
Stems | erect to ascending, usually simple, 2–10(–20) cm, villous, internodes shortened. |
erect, simple or branched from base, 7–28 cm, usually moderately to densely villous-glandular, without eglandular hairs. |
Leaves | all basal or near basal, often rosulate; petiole 0 mm; blade palmately 3-veined, oblanceolate to elliptic-obovate, 7–40 × 4–12 mm, base cuneate to attenuate, margins entire, distally denticulate to dentate, or sharply serrate-dentate, apex acute to obtuse, abaxial surface glabrous, adaxial glabrous or glabrate to sparsely to densely long-villous, eglandular. |
usually cauline or basal persistent; petiole: proximals to medials 7–15 mm, distals 0 mm; blade palmately (3–)5-veined, ovate or broadly ovate to elliptic-ovate or depressed-ovate, (10–)15–30 × 6–15 mm, base truncate to shallowly cuneate, margins shallowly to coarsely dentate, apex acute, surfaces usually moderately to densely villous-glandular, without eglandular hairs. |
Flowers | herkogamous, 1. |
herkogamous, (1–)2–6, axillary from middle to distal nodes, chasmogamous. |
Styles | glabrous. |
glabrous or sparsely hirtellous. |
Corollas | yellow to orange-yellow, usually brown-spotted abaxially, base of each abaxial lobe usually with a larger reddish brown spot, bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular, densely hirsute on abaxial side of opening; tube-throat narrowly campanulate, 15–20 mm, exserted beyond calyx margin; lobes broadly obovate-oblong, apex rounded- or truncate-notched, throat open, palate densely villous, abaxial ridges prominent. |
yellow, throat red-spotted, abaxial limb base with a large red splotch, bilaterally symmetric, bilabiate; tube-throat narrowly cylindric-funnelform, 10–12 mm, exserted 4–5 mm beyond calyx margin; limb abruptly expanded 14–20 mm. |
Fruiting pedicels | 30–110(–130) mm, glabrous or sparsely stipitate-glandular near base. |
15–45 mm, usually moderately to densely villous-glandular, without eglandular hairs. |
Fruiting calyces | tubular-campanulate, weakly or not inflated, 6–8 mm, glabrous. |
sharply nodding, usually densely purple-spotted, broadly campanulate, inflated, sagittally compressed, 9–12 mm, densely hirtellous, sometimes sparsely stipitate-glandular, densely villous at sinuses, throat closing, adaxial lobe ca. 2 times length of others. |
Capsules | included, 6–7 mm. |
included, 6–9 mm. |
Anthers | slightly exserted, margins ciliate, glabrous. |
included, glabrous. |
2n | = 34. |
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Erythranthe primuloides |
Erythranthe marmorata |
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Phenology | Flowering Jun–Aug. | Flowering Mar–May. |
Habitat | Wet meadows, seeps, streamsides. | Habitat unknown, not over serpentine. |
Elevation | 600–3400 m. (2000–11200 ft.) | 100–900 m. (300–3000 ft.) |
Distribution |
AZ; CA; ID; MT; NM; NV; OR; UT; WA
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CA |
Discussion | Flowers in Erythranthe primuloides and E. linearifolia characteristically appear to be scapose, but the scapes are pedicels arising from axils of greatly foreshortened stems. Occasionally in both species the internodes may lengthen somewhat, and the leaves are not so densely clustered at the base of the stems. In northern Klamath, western Deschutes, and eastern Douglas counties, Oregon, an area within the range of typical populations, Erythranthe primuloides has distinctively large corollas (limbs 10–15 mm wide). Apparent clones of large-flowered and smaller-flowered plants sometimes grow in close proximity or even intermixed, appearing as two different entities. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe marmorata is recognized by its erect, taprooted habit and annual duration, villous-glandular vestiture, ovate-petiolate leaves, flowers from middle to distal nodes, long, narrow corolla tube-throat abruptly flaring into a broad limb, abaxial middle corolla lobe with a large red splotch, and fruiting calyces dark-spotted and sharply nodding. The species is known from foothills in Calaveras, Stanislaus, and Tuolumne counties in the Stanislaus River drainage and from Amador County in the Mokelumne River drainage. A collection from Fresno County appears to be somewhat disjunct from the main range, and the plants are more densely villous than characteristic elsewhere, but their identification as E. marmorata otherwise seems secure. Erythranthe marmorata (previously identified as Mimulus whipplei) had been considered extremely rare or even perhaps extinct. See G. L. Nesom (2013d) for citations of recent collections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 389. | FNA vol. 17, p. 416. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus primuloides, M. nevadensis, M. pilosellus, M. primuloides var. minimus, M. primuloides var. pilosellus | Mimulus marmoratus, M. whipplei |
Name authority | (Bentham) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 35. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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