Erythranthe primuloides |
Erythranthe geniculata |
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primrose monkey-flower |
bent-stem monkeyflower, Dudley's monkeyflower |
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Habit | Perennials, rhizomatous or stoloniferous, mat-forming, rhizomes or stolons flagelliform. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | erect to ascending, usually simple, 2–10(–20) cm, villous, internodes shortened. |
ascending to decumbent or prostrate, geniculate at nodes, simple or diffusely branched, 5–60 cm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
Leaves | all basal or near basal, often rosulate; petiole 0 mm; blade palmately 3-veined, oblanceolate to elliptic-obovate, 7–40 × 4–12 mm, base cuneate to attenuate, margins entire, distally denticulate to dentate, or sharply serrate-dentate, apex acute to obtuse, abaxial surface glabrous, adaxial glabrous or glabrate to sparsely to densely long-villous, eglandular. |
basal and cauline, basal usually deciduous by flowering; petiole 2–10(–35) mm; blade pinnately to subpinnately veined, broadly ovate or elliptic-ovate to triangular, 8–35 × 5–30 mm, base cuneate to rounded or subcordate, margins serrate or dentate, teeth 3–10 per side, apex acute to obtuse or rounded, surfaces moderately villous, hairs 0.8–2 mm, multicellular, eglandular, and 0.1–0.3 mm, stipitate-glandular. |
Flowers | herkogamous, 1. |
herkogamous, (1–)6–20, from all or medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow to orange-yellow, usually brown-spotted abaxially, base of each abaxial lobe usually with a larger reddish brown spot, bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular, densely hirsute on abaxial side of opening; tube-throat narrowly campanulate, 15–20 mm, exserted beyond calyx margin; lobes broadly obovate-oblong, apex rounded- or truncate-notched, throat open, palate densely villous, abaxial ridges prominent. |
yellow, without white patches, throat red-spotted, spots concentrated or becoming coalescent into a somewhat discrete splotch at base of each of 3 abaxial lobes and sometimes 2 adaxial, bilaterally symmetric, ± bilabiate; tube-throat cylindric, 9–12 mm, exserted beyond calyx margin; limb expanded 10–18 mm diam. |
Fruiting pedicels | 30–110(–130) mm, glabrous or sparsely stipitate-glandular near base. |
12–26(–55) mm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
Fruiting calyces | tubular-campanulate, weakly or not inflated, 6–8 mm, glabrous. |
red-spotted, campanulate-cylindric, weakly inflated, (5–)6–8 mm, margins distinctly toothed or lobed, sparsely to moderately villous-glandular, ribs shallowly wing-angled, lobes pronounced, erect to spreading or spreading-recurving. |
Capsules | included, 6–7 mm. |
included, 4–6(–7) mm. |
Anthers | slightly exserted, margins ciliate, glabrous. |
included, glabrous. |
2n | = 34. |
= 32. |
Erythranthe primuloides |
Erythranthe geniculata |
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Phenology | Flowering Jun–Aug. | Flowering (Mar–)Apr–Jul. |
Habitat | Wet meadows, seeps, streamsides. | Granite crevices, canyon slopes, talus, crevices in volcanic outcrops, edges of boulders, roadsides, damp sandy soils, sandy water edges, gravelly soils and creek bottoms. |
Elevation | 600–3400 m. (2000–11200 ft.) | 200–900(–1200) m. (700–3000(–3900) ft.) |
Distribution |
AZ; CA; ID; MT; NM; NV; OR; UT; WA
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CA
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Discussion | Flowers in Erythranthe primuloides and E. linearifolia characteristically appear to be scapose, but the scapes are pedicels arising from axils of greatly foreshortened stems. Occasionally in both species the internodes may lengthen somewhat, and the leaves are not so densely clustered at the base of the stems. In northern Klamath, western Deschutes, and eastern Douglas counties, Oregon, an area within the range of typical populations, Erythranthe primuloides has distinctively large corollas (limbs 10–15 mm wide). Apparent clones of large-flowered and smaller-flowered plants sometimes grow in close proximity or even intermixed, appearing as two different entities. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe geniculata is known from an apparently disjunct cluster of populations in Butte, Sutter, and Yuba counties and then from Tuolumne and Stanislaus counties south to Kern County. Erythranthe geniculata, compared to E. floribunda, has larger, chasmogamous, and allogamous flowers. The anther pairs of E. geniculata are at different levels, and the stigma is slightly above the adaxial anther pair; in E. floribunda both anther pairs and the stigma are at the same level. Erythranthe arenaria, E. geniculata, and E. norrisii constitute a group of apparently closely related species within sect. Mimulosma endemic along the Sierra Nevada. All have ovate-petiolate leaves (only the basal ones are sometimes ovate in E. arenaria) with pinnate to subpinnate venation. The more widespread E. floribunda, which is part of the above group, also is similar, but all three endemics have larger corollas with the tube exserted at greater length beyond the calyx margin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 389. | FNA vol. 17, p. 404. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus primuloides, M. nevadensis, M. pilosellus, M. primuloides var. minimus, M. primuloides var. pilosellus | Mimulus geniculatus, M. dudleyi, M. floribundus var. geniculatus |
Name authority | (Bentham) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 35. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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