Erythranthe primuloides |
Erythranthe arvensis |
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primrose monkey-flower |
field monkey-flower, villous-bract monkeyflower, western monkey-flower |
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Habit | Perennials, rhizomatous or stoloniferous, mat-forming, rhizomes or stolons flagelliform. | Annuals, taprooted or fibrous-rooted, sometimes rooting at proximal cauline nodes if decumbent. |
Stems | erect to ascending, usually simple, 2–10(–20) cm, villous, internodes shortened. |
erect to decumbent-ascending, simple or branched from proximal to medial nodes, usually 4-angled, fistulose to very narrow, 5–70 cm, glabrous, sometimes minutely hirtellous in inflorescence, hairs deflexed, eglandular. |
Leaves | all basal or near basal, often rosulate; petiole 0 mm; blade palmately 3-veined, oblanceolate to elliptic-obovate, 7–40 × 4–12 mm, base cuneate to attenuate, margins entire, distally denticulate to dentate, or sharply serrate-dentate, apex acute to obtuse, abaxial surface glabrous, adaxial glabrous or glabrate to sparsely to densely long-villous, eglandular. |
basal and cauline or basal not persistent, often largest at mid stem or above, reduced in size distally; petiole 3–20(–90) mm, distals 0 mm; blade palmately 3–5-veined, ovate to orbicular, orbicular-ovate, oblong-ovate, or (middle and distal cauline) broadly orbicular to depressed-ovate or nearly reniform, (5–)10–35(–45) × 6–26(–50) mm, distal closely paired, auriculate-subclasping, base rounded to truncate, subcordate, or shallowly cordate, margins denticulate or subentire to distinctly dentate, on larger plants proximal characteristically lacerate-lobed to pinnatifid at margin base, apex rounded, surfaces glabrous except for bracts densely villous abaxially, sometimes also adaxially, hairs long, sometimes vitreous, flattened, eglandular, multicellular. |
Flowers | herkogamous, 1. |
plesiogamous, 3–8(–16), from remote distal nodes, chasmogamous or cleistogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow to orange-yellow, usually brown-spotted abaxially, base of each abaxial lobe usually with a larger reddish brown spot, bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular, densely hirsute on abaxial side of opening; tube-throat narrowly campanulate, 15–20 mm, exserted beyond calyx margin; lobes broadly obovate-oblong, apex rounded- or truncate-notched, throat open, palate densely villous, abaxial ridges prominent. |
yellow, usually red-spotted, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat cylindric-funnelform, (7–)8–12 mm, exserted (0–)1–2(–3) mm beyond calyx margin; limb expanded 5–10 mm. |
Fruiting pedicels | 30–110(–130) mm, glabrous or sparsely stipitate-glandular near base. |
5–40(–90) mm, longer than subtending leaves, glabrous. |
Fruiting calyces | tubular-campanulate, weakly or not inflated, 6–8 mm, glabrous. |
red-dotted or not, ovate-campanulate, inflated, sagittally compressed, (7–)9–14 mm, minutely hirtellous, throat closing or not, remaining open, lobes upcurving weakly, adaxial lobe not distinctly longer than abaxial, not falcate. |
Capsules | included, 6–7 mm. |
included, stipitate, (5–)6–7 mm. |
Anthers | slightly exserted, margins ciliate, glabrous. |
included, glabrous. |
2n | = 34. |
= 28. |
Erythranthe primuloides |
Erythranthe arvensis |
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Phenology | Flowering Jun–Aug. | Flowering Apr–Jun(–Jul). |
Habitat | Wet meadows, seeps, streamsides. | Hills, ridges, clay banks, stream banks, moist woods. |
Elevation | 600–3400 m. (2000–11200 ft.) | 30–1900(–2300) m. (100–6200(–7500) ft.) |
Distribution |
AZ; CA; ID; MT; NM; NV; OR; UT; WA
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CA; ID; MT; NV; OR; UT; WA; WY; BC; Mexico (Baja California)
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Discussion | Flowers in Erythranthe primuloides and E. linearifolia characteristically appear to be scapose, but the scapes are pedicels arising from axils of greatly foreshortened stems. Occasionally in both species the internodes may lengthen somewhat, and the leaves are not so densely clustered at the base of the stems. In northern Klamath, western Deschutes, and eastern Douglas counties, Oregon, an area within the range of typical populations, Erythranthe primuloides has distinctively large corollas (limbs 10–15 mm wide). Apparent clones of large-flowered and smaller-flowered plants sometimes grow in close proximity or even intermixed, appearing as two different entities. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe arvensis usually is easily recognized, characterized by its annual duration (but commonly rooting at proximal cauline nodes, suggestive of a rhizomatous habit), glabrous stems with nodes relatively few and remotely spaced, depressed-ovate leaves with margins often sublyrate (lacerate-lobed to subpinnatifid) at the base, distal leaves and bracts densely villous with vitreous eglandular hairs, other leaves glabrous, and corollas varying in size from relatively small but perhaps chasmogamous (the type of Mimulus arvensis) to even smaller (cleistogamous; the type of M. micranthus). The breeding system is consistently autogamous. The relatively short and even-sized calyx lobes that do not turn upward to close the orifice have been considered diagnostic of E. arvensis. This feature is evident in some plants, but others (perhaps reflecting gene flow from other species) have a longer adaxial calyx lobe and abaxial lobes that turn upward variably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 389. | FNA vol. 17, p. 420. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus primuloides, M. nevadensis, M. pilosellus, M. primuloides var. minimus, M. primuloides var. pilosellus | Mimulus arvensis, M. guttatus subsp. arvensis, M. guttatus var. arvensis, M. guttatus subsp. micranthus, M. langsdorffii var. arvensis, M. longulus, M. micranthus |
Name authority | (Bentham) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 35. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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