Erythranthe primuloides |
Erythranthe arenicola |
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primrose monkey-flower |
beach monkeyflower |
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Habit | Perennials, rhizomatous or stoloniferous, mat-forming, rhizomes or stolons flagelliform. | Annuals, fibrous-rooted or slender-taprooted, rarely rooting at nodes. |
Stems | erect to ascending, usually simple, 2–10(–20) cm, villous, internodes shortened. |
erect, rarely prostrate to prostrate-ascending, few-branched, 3–17 cm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Leaves | all basal or near basal, often rosulate; petiole 0 mm; blade palmately 3-veined, oblanceolate to elliptic-obovate, 7–40 × 4–12 mm, base cuneate to attenuate, margins entire, distally denticulate to dentate, or sharply serrate-dentate, apex acute to obtuse, abaxial surface glabrous, adaxial glabrous or glabrate to sparsely to densely long-villous, eglandular. |
basal and cauline; petiole: basal 2–8 mm or mid and distals absent; blade palmately 3–5-veined, suborbicular to broadly ovate or depressed-ovate, 5–17 × 6–15 mm, base truncate or truncate-cuneate to subcordate, margins subentire or crenulate, apex rounded to obtuse, surfaces moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Flowers | herkogamous, 1. |
herkogamous, 1–6, at distal nodes, chasmogamous. |
Styles | glabrous. |
hirtellous. |
Corollas | yellow to orange-yellow, usually brown-spotted abaxially, base of each abaxial lobe usually with a larger reddish brown spot, bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular, densely hirsute on abaxial side of opening; tube-throat narrowly campanulate, 15–20 mm, exserted beyond calyx margin; lobes broadly obovate-oblong, apex rounded- or truncate-notched, throat open, palate densely villous, abaxial ridges prominent. |
yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 11–20 mm, exserted 4–8 mm beyond calyx margin; limb expanded 10–18 mm. |
Fruiting pedicels | 30–110(–130) mm, glabrous or sparsely stipitate-glandular near base. |
9–17 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Fruiting calyces | tubular-campanulate, weakly or not inflated, 6–8 mm, glabrous. |
nodding, ovoid-campanulate, inflated, sagittally compressed, 9–16 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular, throat closing. |
Capsules | included, 6–7 mm. |
included, 5–12 mm. |
Anthers | slightly exserted, margins ciliate, glabrous. |
included, glabrous. |
2n | = 34. |
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Erythranthe primuloides |
Erythranthe arenicola |
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Phenology | Flowering Jun–Aug. | Flowering Apr–Aug. |
Habitat | Wet meadows, seeps, streamsides. | Sandy beaches, especially in moist hollows among dunes, sea cliff bases, chaparral near beaches, mudstone outcrops. |
Elevation | 600–3400 m. (2000–11200 ft.) | 0–100 m. (0–300 ft.) |
Distribution |
AZ; CA; ID; MT; NM; NV; OR; UT; WA
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CA |
Discussion | Flowers in Erythranthe primuloides and E. linearifolia characteristically appear to be scapose, but the scapes are pedicels arising from axils of greatly foreshortened stems. Occasionally in both species the internodes may lengthen somewhat, and the leaves are not so densely clustered at the base of the stems. In northern Klamath, western Deschutes, and eastern Douglas counties, Oregon, an area within the range of typical populations, Erythranthe primuloides has distinctively large corollas (limbs 10–15 mm wide). Apparent clones of large-flowered and smaller-flowered plants sometimes grow in close proximity or even intermixed, appearing as two different entities. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
F. W. Pennell (1947, 1951) considered Erythranthe arenicola an endemic of Monterey County, but plants from adjacent San Luis Obispo and Santa Cruz counties also belong here. Most of the localities are at seaside, but some are more than a mile inland. Erythranthe arenicola is hypothesized here to be a derivative of E. guttata or E. grandis, retaining the herkogamous breeding system of its putative ancestor but reduced in size and duration. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 389. | FNA vol. 17, p. 412. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus primuloides, M. nevadensis, M. pilosellus, M. primuloides var. minimus, M. primuloides var. pilosellus | Mimulus guttatus subsp. arenicola |
Name authority | (Bentham) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 35. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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