Erythranthe patula |
Phrymaceae |
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stalk-leaf monkey-flower |
lop-seed family, monkeyflower family |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Herbs, subshrubs, or shrubs, annual or perennial, aquatic or terrestrial, sometimes fleshy, autotrophic. | ||||||||||||||||||||||||
Stems | erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
erect or ascending to prostrate, 4-angled, sometimes winged. |
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Leaves | cauline, basal not persistent; petiole (5–)8–25 mm; blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
deciduous or persistent, basal and cauline or all cauline, rarely subrosulate or rosulate (Erythranthe), opposite, or alternate distally, simple; stipules absent; petiole present or absent; blade fleshy, semi-fleshy, or not, not leathery, margins entire or toothed. |
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Inflorescences | terminal and axillary racemes or flowers solitary (Glossostigma, some Erythranthe, some annual plants); flowers erect to nodding or strongly reflexed and appressed to inflorescence axis (Phryma). |
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Flowers | herkogamous, 1–10, from proximal to distal nodes. |
bisexual, perianth and androecium hypogynous; sepals (3 or)4 or 5, proximally connate, calyx radially or bilaterally symmetric; petals 3–5, proximally connate, corolla bilaterally symmetric, rarely nearly radially in reduced forms, strongly to weakly bilabiate, rarely nearly regular, salverform to tubular-funnelform, funnelform, campanulate, or compressed; stamens (2–)4, adnate to corolla, didynamous [both pairs of equal length in autogamous forms], staminode 0; pistil 1, 2-carpellate, ovary superior, (1 or)2-locular, placentation axile, basal (Phryma), or parietal (Diplacus, Mimetanthe); ovules anatropous or orthotropous (Phryma), unitegmic, tenuinucellate; style 1; stigma 1, 2-lobed. |
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Styles | glabrous. |
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Corollas | yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate; tube-throat funnelform, 7–8 mm, exserted beyond calyx margin; lobes oblong, apex rounded to truncate. |
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Fruiting pedicels | 10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
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Fruiting calyces | tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect. |
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Fruits | capsules, dehiscence loculicidal [septicidal or irregular], or achenes [berry]. |
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Capsules | included, 4–6 mm. |
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Seeds | 1–2000, yellowish brown or brown, narrowly ellipsoid, slightly flattened bilaterally; embryo straight, endosperm sparse. |
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Anthers | included, glabrous. |
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2n | = 32. |
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Erythranthe patula |
Phrymaceae |
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Phenology | Flowering Apr–May(–Aug). | |||||||||||||||||||||||||
Habitat | Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices. | |||||||||||||||||||||||||
Elevation | 200–1900(–2900) m. (700–6200(–9500) ft.) | |||||||||||||||||||||||||
Distribution |
ID; MT; OR; WA; WY; AB; BC
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North America; Mexico; Central America; w South America (primarily Andean); s Asia (India); se Asia; e Africa; Indian Ocean Islands (Madagascar); Pacific Islands (New Zealand); Australia [Introduced in Europe, s Africa] |
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Discussion | Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 13, species ca. 200 (6 genera, 139 species in the flora). Over one half of the species in Phrymaceae are members of Diplacus and Erythranthe and together include over 160 species; all other genera each have seven or fewer species. Until recently, Phrymaceae consisted only of Phryma leptostachya, a taxonomically isolated species of eastern North America and eastern Asia. Molecular studies have established a relationship not with the Verbenaceae, as was earlier postulated (see H. L. Whipple 1972; R. Venkata Ramana et al. 2000), but rather with Mimulus and other genera, suggesting that Phrymaceae should be enlarged. The sequence of genera in Phrymaceae here follows the phylogeny proposed by P. M. Beardsley and R. G. Olmstead (2002). One of the major lineages of Phrymaceae is primarily a Southern Hemisphere group ranging from Australia and New Zealand to southeastern and south Asia (India), Madagascar, and South Africa. Mimulus in the narrow sense, including the two endemic North American species, is part of this group, which includes 24 species in seven genera. The largest major lineage includes 158 species in five genera from North America, South America, and southeast Asia. This lineage includes two genera from Mexico and Central America: Hemichaena Bentham, which is sister to the North American Diplacus and Mimetanthe, and Leucocarpus D. Don, which is sister to the American and Asian Erythranthe. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 397. | FNA vol. 17, p. 365. | ||||||||||||||||||||||||
Parent taxa | Phrymaceae > Erythranthe | |||||||||||||||||||||||||
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Synonyms | Mimulus patulus | |||||||||||||||||||||||||
Name authority | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) | Schauer | ||||||||||||||||||||||||
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