Erythranthe patula |
Erythranthe scouleri |
|
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stalk-leaf monkey-flower |
Columbia River monkey-flower, Scouler's monkey-flower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Perennials, rhizomatous, producing long, sparsely leafy runners from basal nodes. |
Stems | erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
erect, simple or few-branched, 15–80 cm, glabrous. |
Leaves | cauline, basal not persistent; petiole (5–)8–25 mm; blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
cauline, basal usually not persistent; petiole: proximals and medial 10–25 mm (gradually merging into blade), distals 0 mm; blade palmately (3–)5–7-veined to subpinnate, (proximal to medial) oblong-elliptic to oblong-lanceolate, 25–60 × 8–18 mm, usually 3–4 times longer than wide, base attenuate, margins evenly, shallowly dentate or crenate to mucronate or mucronulate, teeth 10–20 per side, sometimes more deeply toothed at base, apex obtuse to acute, surfaces glabrous. |
Flowers | herkogamous, 1–10, from proximal to distal nodes. |
herkogamous, (1 or)2–8, from distal nodes. |
Styles | glabrous. |
minutely, prominently hirsutulous to villosulous. |
Corollas | yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate; tube-throat funnelform, 7–8 mm, exserted beyond calyx margin; lobes oblong, apex rounded to truncate. |
yellow, without red markings, bilaterally symmetric, bilabiate; tube-throat funnelform, 20–24 mm, exserted 10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
Fruiting pedicels | 10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
20–25 mm, glabrous. |
Fruiting calyces | tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect. |
ovoid, inflated, sagittally compressed, 13–14 mm, glabrous, throat closing. |
Capsules | included, 4–6 mm. |
unknown. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe patula |
Erythranthe scouleri |
|
Phenology | Flowering Apr–May(–Aug). | Flowering May–Aug. |
Habitat | Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices. | Salt marshes, river banks. |
Elevation | 200–1900(–2900) m. (700–6200(–9500) ft.) | 0–20 m. (0–100 ft.) |
Distribution |
ID; MT; OR; WA; WY; AB; BC
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OR; WA |
Discussion | Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe scouleri is distinctive in its oblong-elliptic leaves with long-tapering bases and closely toothed margins, completely glabrous vestiture, long, leafy runners from basal cauline nodes, large corollas with broad limbs, and prominently hairy styles. Several features suggest a close relationship to E. decora, particularly its very large corollas, hairy styles, closely toothed leaf margins, tall, simple, and erect stems, numerous runners, and its geographic range. All collections apparently have been made near the mouth of the Columbia River in Clatsop and Columbia counties, Oregon (G. L. Nesom 2013d). Recent observations (Alexander John Wright, pers. comm.; photos http://www.inaturalist.org/observations/3537008) indicate that it also occurs in Wahkiakum County, Washington, in the Julia Butler Hansen Refuge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 397. | FNA vol. 17, p. 415. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus patulus | Mimulus scouleri, M. guttatus subsp. scouleri |
Name authority | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) | (Hooker) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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