FNA: Erythranthe patula vs. Erythranthe cordata

	Erythranthe patula	Erythranthe cordata
	stalk-leaf monkey-flower	tinytooter monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Annuals, fibrous-rooted, sometimes producing leafy runners from basal nodes, stems often rooting at proximal nodes and appearing rhizomelike.
Stems	erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.	usually erect, usually simple, usually fistulose, 12–40(–100) cm, sparsely stipitate-glandular, hairs fine, gland-tipped.
Leaves	cauline, basal not persistent; petiole (5–)8–25 mm; blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.	basal and cauline, basal persistent; petiole: basal and proximals 6–20(–40) mm, midcauline to distals 0 mm; blade not connate, palmately 3–5(–7)-veined, orbicular to broadly elliptic-ovate or oblong-elliptic, cauline becoming broadly ovate to narrowly reniform, basal and mid cauline 15–30(–50) mm, gradually reduced in size distally to 6 mm, basal largest, distal closely paired, auriculate-subclasping, base cuneate to truncate or shallowly cordate, margins shallowly, evenly to unevenly dentate, apex obtuse to rounded, surfaces glabrous.
Flowers	herkogamous, 1–10, from proximal to distal nodes.	plesiogamous, (5–)10–16, at distal nodes, in bracteate racemes, chasmogamous or cleistogamous.
Styles	glabrous.	glabrous.
Corollas	yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate; tube-throat funnelform, 7–8 mm, exserted beyond calyx margin; lobes oblong, apex rounded to truncate.	yellow, red-spotted, abaxial limb deeper yellow, weakly bilaterally or radially symmetric, weakly bilabiate or regular; tube-throat sometimes tubular and not opening (cleistogamous), 8–14 mm, exserted 1–3 mm beyond calyx margin; limb not expanded or expanded 9–14 mm.
Fruiting pedicels	10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.	10–30(–45) mm, longer than subtending leaves, minutely stipitate-glandular.
Fruiting calyces	tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect.	nodding 45–90º, not red-dotted, broadly elliptic-ovoid, inflated, sagittally compressed, (8–)14–18(–20) mm, glabrous or sparsely stipitate-glandular to hirsutulous, sometimes mixed glandular-hirsutulous, throat closing, adaxial lobe not distinctly longer than abaxial, not falcate.
Capsules	included, 4–6 mm.	included, stipitate, 5–7 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 32.	= 60.

	Erythranthe patula	Erythranthe cordata
Phenology	Flowering Apr–May(–Aug).	Flowering (Jan–)Mar–Jun(–Nov).
Habitat	Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices.	Springs, seeps, stream edges, muddy banks, flood plains, marshes and swamps, wash bottoms, wet depressions, wet places among boulders.
Elevation	200–1900(–2900) m. (700–6200(–9500) ft.)	(600–)800–2400(–3000) m. ((2000–)2600–7900(–9800) ft.)
Distribution	ID; MT; OR; WA; WY; AB; BC [WildflowerSearch map]	AZ; CA; CO; NM; NV; TX; UT [WildflowerSearch map]
Discussion	Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe cordata is characterized by its fibrous-rooted habit (annual in duration, without rhizomes but commonly rooting at the proximal nodes), short corollas and autogamous reproduction (anthers and stigma at the same level), closed calyces, sparsely villous-glandular vestiture (lacking hirtellous, eglandular hairs), and stems commonly fistulose in larger plants. The short corollas and other features of autogamous reproduction of E. cordata are diagnostic and prominent. Plants of E. cordata are highly variable in size, from tiny fibrous-rooted plants with nearly filiform stems to much larger individuals with fistulose stems rooting at proximal nodes. Erythranthe cordata and E. nasuta are sympatric in Arizona, southeastern New Mexico, and southern Utah, and small plants of each species may be similar in aspect, both with cleistogamous flowers and reduced vestiture. Erythranthe nasuta can be recognized by its distal and bracteal leaves with hirtellous to hirsutulous adaxial surfaces; a 10/x lens usually is required to see this feature, and it sometimes is most obvious around the leaf margins. The common name of Erythranthe cordata alludes to a fancied resemblance of the corollas to the horn of a diminutive trumpet. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 397.	FNA vol. 17, p. 422.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus patulus	Mimulus cordatus, M. maguirei
Name authority	(Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012)	(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)
Web links	Local floras: BC, OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe cordata

stalk-leaf monkey-flower

tinytooter monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Annuals, fibrous-rooted, sometimes producing leafy runners from basal nodes, stems often rooting at proximal nodes and appearing rhizomelike.

Stems

erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.

usually erect, usually simple, usually fistulose, 12–40(–100) cm, sparsely stipitate-glandular, hairs fine, gland-tipped.

Leaves

cauline, basal not persistent;

petiole (5–)8–25 mm;

blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.

basal and cauline, basal persistent;

petiole: basal and proximals 6–20(–40) mm, midcauline to distals 0 mm;

blade not connate, palmately 3–5(–7)-veined, orbicular to broadly elliptic-ovate or oblong-elliptic, cauline becoming broadly ovate to narrowly reniform, basal and mid cauline 15–30(–50) mm, gradually reduced in size distally to 6 mm, basal largest, distal closely paired, auriculate-subclasping, base cuneate to truncate or shallowly cordate, margins shallowly, evenly to unevenly dentate, apex obtuse to rounded, surfaces glabrous.

Flowers

herkogamous, 1–10, from proximal to distal nodes.

plesiogamous, (5–)10–16, at distal nodes, in bracteate racemes, chasmogamous or cleistogamous.

Styles

glabrous.

Corollas

yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate;

tube-throat funnelform, 7–8 mm, exserted beyond calyx margin;

lobes oblong, apex rounded to truncate.

yellow, red-spotted, abaxial limb deeper yellow, weakly bilaterally or radially symmetric, weakly bilabiate or regular;

tube-throat sometimes tubular and not opening (cleistogamous), 8–14 mm, exserted 1–3 mm beyond calyx margin;

limb not expanded or expanded 9–14 mm.

Fruiting pedicels

10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.

10–30(–45) mm, longer than subtending leaves, minutely stipitate-glandular.

Fruiting calyces

tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect.

nodding 45–90º, not red-dotted, broadly elliptic-ovoid, inflated, sagittally compressed, (8–)14–18(–20) mm, glabrous or sparsely stipitate-glandular to hirsutulous, sometimes mixed glandular-hirsutulous, throat closing, adaxial lobe not distinctly longer than abaxial, not falcate.

Capsules

included, 4–6 mm.

included, stipitate, 5–7 mm.

Anthers

included, glabrous.

= 32.

= 60.

Erythranthe patula

Erythranthe cordata

Phenology

Flowering Apr–May(–Aug).

Flowering (Jan–)Mar–Jun(–Nov).

Habitat

Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices.

Springs, seeps, stream edges, muddy banks, flood plains, marshes and swamps, wash bottoms, wet depressions, wet places among boulders.

Elevation

200–1900(–2900) m. (700–6200(–9500) ft.)

(600–)800–2400(–3000) m. ((2000–)2600–7900(–9800) ft.)

Distribution

ID; MT; OR; WA; WY; AB; BC

[WildflowerSearch map]

AZ; CA; CO; NM; NV; TX; UT

[WildflowerSearch map]

Discussion

Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe cordata is characterized by its fibrous-rooted habit (annual in duration, without rhizomes but commonly rooting at the proximal nodes), short corollas and autogamous reproduction (anthers and stigma at the same level), closed calyces, sparsely villous-glandular vestiture (lacking hirtellous, eglandular hairs), and stems commonly fistulose in larger plants. The short corollas and other features of autogamous reproduction of E. cordata are diagnostic and prominent. Plants of E. cordata are highly variable in size, from tiny fibrous-rooted plants with nearly filiform stems to much larger individuals with fistulose stems rooting at proximal nodes.

Erythranthe cordata and E. nasuta are sympatric in Arizona, southeastern New Mexico, and southern Utah, and small plants of each species may be similar in aspect, both with cleistogamous flowers and reduced vestiture. Erythranthe nasuta can be recognized by its distal and bracteal leaves with hirtellous to hirsutulous adaxial surfaces; a 10/x lens usually is required to see this feature, and it sometimes is most obvious around the leaf margins.

The common name of Erythranthe cordata alludes to a fancied resemblance of the corollas to the horn of a diminutive trumpet.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 397.

FNA vol. 17, p. 422.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus patulus

Mimulus cordatus, M. maguirei

Name authority

(Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012)

(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Erythranthe patula

Erythranthe cordata

Erythranthe patula

Erythranthe cordata