FNA: Erythranthe patula vs. Erythranthe calcicola

	Erythranthe patula	Erythranthe calcicola
	stalk-leaf monkey-flower	limestone monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Annuals, taprooted.
Stems	erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.	erect, simple or branched from basal nodes, 2–10(–15) cm, sparsely glandular-pubescent, internodes elongate, distinct.
Leaves	cauline, basal not persistent; petiole (5–)8–25 mm; blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.	cauline, basal not persistent; petiole 0–1 mm; blade palmately 3-veined (in broader ones), lanceolate to ovate, 3–25 × 2–8(–10) mm, base attenuate, margins entire, sometimes toothed, apex acute to obtuse, surfaces sparsely glandular-pubescent.
Flowers	herkogamous, 1–10, from proximal to distal nodes.	herkogamous, sometimes plesiogamous, 1–16, from distal or medial to distal nodes.
Styles	glabrous.	glabrous.
Corollas	yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate; tube-throat funnelform, 7–8 mm, exserted beyond calyx margin; lobes oblong, apex rounded to truncate.	yellow or white and throat yellow, throat and limb red-spotted, bilaterally symmetric, weakly bilabiate; tube-throat cylindric, 6–13 mm, exserted beyond calyx margin; limb expanded 3–7(–9) mm, lobes notched, abaxial limb sparsely bearded or glabrous.
Fruiting pedicels	10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.	(3–)5–20 mm.
Fruiting calyces	tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect.	campanulate to widely cylindric, 5–7 mm, margins distinctly toothed or lobed, sparsely glandular-pubescent, ribs strongly angled, lobes pronounced, erect.
Capsules	included, 4–6 mm.	included, 4–6 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 32.

	Erythranthe patula	Erythranthe calcicola
Phenology	Flowering Apr–May(–Aug).	Flowering Apr–Jun.
Habitat	Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices.	Creosote bush and Joshua tree woodlands, usually on carbonate substrate, primarily on talus slopes.
Elevation	200–1900(–2900) m. (700–6200(–9500) ft.)	900–2200 m. (3000–7200 ft.)
Distribution	ID; MT; OR; WA; WY; AB; BC [WildflowerSearch map]	CA; NV
Discussion	Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe calcicola was previously included in the broader concept of E. montioides; the former can be distinguished by its glandular herbage, broader lanceolate to ovate leaves, strongly angled calyces, and smaller flowers. The species is restricted to the northern Mojave Desert and southwestern Great Basin and occurs primarily on talus slopes on substrates derived from carbonate rock. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 397.	FNA vol. 17, p. 383.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus patulus
Name authority	(Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012)	N. S. Fraga & D. A. York: Aliso 30: 54, figs. 12–16. (2012)
Web links	Local floras: BC, OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe calcicola

stalk-leaf monkey-flower

limestone monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Annuals, taprooted.

Stems

erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.

erect, simple or branched from basal nodes, 2–10(–15) cm, sparsely glandular-pubescent, internodes elongate, distinct.

Leaves

cauline, basal not persistent;

petiole (5–)8–25 mm;

blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.

cauline, basal not persistent;

petiole 0–1 mm;

blade palmately 3-veined (in broader ones), lanceolate to ovate, 3–25 × 2–8(–10) mm, base attenuate, margins entire, sometimes toothed, apex acute to obtuse, surfaces sparsely glandular-pubescent.

Flowers

herkogamous, 1–10, from proximal to distal nodes.

herkogamous, sometimes plesiogamous, 1–16, from distal or medial to distal nodes.

Styles

glabrous.

Corollas

yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate;

tube-throat funnelform, 7–8 mm, exserted beyond calyx margin;

lobes oblong, apex rounded to truncate.

yellow or white and throat yellow, throat and limb red-spotted, bilaterally symmetric, weakly bilabiate;

tube-throat cylindric, 6–13 mm, exserted beyond calyx margin;

limb expanded 3–7(–9) mm, lobes notched, abaxial limb sparsely bearded or glabrous.

Fruiting pedicels

10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.

(3–)5–20 mm.

Fruiting calyces

tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect.

campanulate to widely cylindric, 5–7 mm, margins distinctly toothed or lobed, sparsely glandular-pubescent, ribs strongly angled, lobes pronounced, erect.

Capsules

included, 4–6 mm.

Anthers

included, glabrous.

= 32.

Erythranthe patula

Erythranthe calcicola

Phenology

Flowering Apr–May(–Aug).

Flowering Apr–Jun.

Habitat

Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices.

Creosote bush and Joshua tree woodlands, usually on carbonate substrate, primarily on talus slopes.

Elevation

200–1900(–2900) m. (700–6200(–9500) ft.)

900–2200 m. (3000–7200 ft.)

Distribution

ID; MT; OR; WA; WY; AB; BC

[WildflowerSearch map]

CA; NV

Discussion

Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe calcicola was previously included in the broader concept of E. montioides; the former can be distinguished by its glandular herbage, broader lanceolate to ovate leaves, strongly angled calyces, and smaller flowers. The species is restricted to the northern Mojave Desert and southwestern Great Basin and occurs primarily on talus slopes on substrates derived from carbonate rock.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 397.

FNA vol. 17, p. 383.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus patulus

Name authority

(Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012)

N. S. Fraga & D. A. York: Aliso 30: 54, figs. 12–16. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Erythranthe patula

Erythranthe calcicola

Erythranthe patula

Erythranthe calcicola