Erythranthe patula |
Erythranthe arvensis |
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stalk-leaf monkey-flower |
field monkey-flower, villous-bract monkeyflower, western monkey-flower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, taprooted or fibrous-rooted, sometimes rooting at proximal cauline nodes if decumbent. |
Stems | erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
erect to decumbent-ascending, simple or branched from proximal to medial nodes, usually 4-angled, fistulose to very narrow, 5–70 cm, glabrous, sometimes minutely hirtellous in inflorescence, hairs deflexed, eglandular. |
Leaves | cauline, basal not persistent; petiole (5–)8–25 mm; blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
basal and cauline or basal not persistent, often largest at mid stem or above, reduced in size distally; petiole 3–20(–90) mm, distals 0 mm; blade palmately 3–5-veined, ovate to orbicular, orbicular-ovate, oblong-ovate, or (middle and distal cauline) broadly orbicular to depressed-ovate or nearly reniform, (5–)10–35(–45) × 6–26(–50) mm, distal closely paired, auriculate-subclasping, base rounded to truncate, subcordate, or shallowly cordate, margins denticulate or subentire to distinctly dentate, on larger plants proximal characteristically lacerate-lobed to pinnatifid at margin base, apex rounded, surfaces glabrous except for bracts densely villous abaxially, sometimes also adaxially, hairs long, sometimes vitreous, flattened, eglandular, multicellular. |
Flowers | herkogamous, 1–10, from proximal to distal nodes. |
plesiogamous, 3–8(–16), from remote distal nodes, chasmogamous or cleistogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate; tube-throat funnelform, 7–8 mm, exserted beyond calyx margin; lobes oblong, apex rounded to truncate. |
yellow, usually red-spotted, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat cylindric-funnelform, (7–)8–12 mm, exserted (0–)1–2(–3) mm beyond calyx margin; limb expanded 5–10 mm. |
Fruiting pedicels | 10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
5–40(–90) mm, longer than subtending leaves, glabrous. |
Fruiting calyces | tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect. |
red-dotted or not, ovate-campanulate, inflated, sagittally compressed, (7–)9–14 mm, minutely hirtellous, throat closing or not, remaining open, lobes upcurving weakly, adaxial lobe not distinctly longer than abaxial, not falcate. |
Capsules | included, 4–6 mm. |
included, stipitate, (5–)6–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 28. |
Erythranthe patula |
Erythranthe arvensis |
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Phenology | Flowering Apr–May(–Aug). | Flowering Apr–Jun(–Jul). |
Habitat | Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices. | Hills, ridges, clay banks, stream banks, moist woods. |
Elevation | 200–1900(–2900) m. (700–6200(–9500) ft.) | 30–1900(–2300) m. (100–6200(–7500) ft.) |
Distribution |
ID; MT; OR; WA; WY; AB; BC
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CA; ID; MT; NV; OR; UT; WA; WY; BC; Mexico (Baja California)
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Discussion | Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe arvensis usually is easily recognized, characterized by its annual duration (but commonly rooting at proximal cauline nodes, suggestive of a rhizomatous habit), glabrous stems with nodes relatively few and remotely spaced, depressed-ovate leaves with margins often sublyrate (lacerate-lobed to subpinnatifid) at the base, distal leaves and bracts densely villous with vitreous eglandular hairs, other leaves glabrous, and corollas varying in size from relatively small but perhaps chasmogamous (the type of Mimulus arvensis) to even smaller (cleistogamous; the type of M. micranthus). The breeding system is consistently autogamous. The relatively short and even-sized calyx lobes that do not turn upward to close the orifice have been considered diagnostic of E. arvensis. This feature is evident in some plants, but others (perhaps reflecting gene flow from other species) have a longer adaxial calyx lobe and abaxial lobes that turn upward variably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 397. | FNA vol. 17, p. 420. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus patulus | Mimulus arvensis, M. guttatus subsp. arvensis, M. guttatus var. arvensis, M. guttatus subsp. micranthus, M. langsdorffii var. arvensis, M. longulus, M. micranthus |
Name authority | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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