Erythranthe patula |
Erythranthe arenicola |
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stalk-leaf monkey-flower |
beach monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, fibrous-rooted or slender-taprooted, rarely rooting at nodes. |
Stems | erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
erect, rarely prostrate to prostrate-ascending, few-branched, 3–17 cm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Leaves | cauline, basal not persistent; petiole (5–)8–25 mm; blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
basal and cauline; petiole: basal 2–8 mm or mid and distals absent; blade palmately 3–5-veined, suborbicular to broadly ovate or depressed-ovate, 5–17 × 6–15 mm, base truncate or truncate-cuneate to subcordate, margins subentire or crenulate, apex rounded to obtuse, surfaces moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Flowers | herkogamous, 1–10, from proximal to distal nodes. |
herkogamous, 1–6, at distal nodes, chasmogamous. |
Styles | glabrous. |
hirtellous. |
Corollas | yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate; tube-throat funnelform, 7–8 mm, exserted beyond calyx margin; lobes oblong, apex rounded to truncate. |
yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 11–20 mm, exserted 4–8 mm beyond calyx margin; limb expanded 10–18 mm. |
Fruiting pedicels | 10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped. |
9–17 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Fruiting calyces | tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect. |
nodding, ovoid-campanulate, inflated, sagittally compressed, 9–16 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular, throat closing. |
Capsules | included, 4–6 mm. |
included, 5–12 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe patula |
Erythranthe arenicola |
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Phenology | Flowering Apr–May(–Aug). | Flowering Apr–Aug. |
Habitat | Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices. | Sandy beaches, especially in moist hollows among dunes, sea cliff bases, chaparral near beaches, mudstone outcrops. |
Elevation | 200–1900(–2900) m. (700–6200(–9500) ft.) | 0–100 m. (0–300 ft.) |
Distribution |
ID; MT; OR; WA; WY; AB; BC
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CA |
Discussion | Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
F. W. Pennell (1947, 1951) considered Erythranthe arenicola an endemic of Monterey County, but plants from adjacent San Luis Obispo and Santa Cruz counties also belong here. Most of the localities are at seaside, but some are more than a mile inland. Erythranthe arenicola is hypothesized here to be a derivative of E. guttata or E. grandis, retaining the herkogamous breeding system of its putative ancestor but reduced in size and duration. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 397. | FNA vol. 17, p. 412. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus patulus | Mimulus guttatus subsp. arenicola |
Name authority | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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