FNA: Erythranthe patula vs. Erythranthe arenaria

	Erythranthe patula	Erythranthe arenaria
	stalk-leaf monkey-flower	sand-loving monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Annuals, fibrous-rooted or filiform-taprooted.
Stems	erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.	erect to ascending, straight or geniculate at nodes, simple or branched, 5–20 cm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped.
Leaves	cauline, basal not persistent; petiole (5–)8–25 mm; blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.	basal and cauline; petiole 0 mm or proximals 1–3(–5) mm; blade 1-veined or palmately 3-veined, elliptic to narrowly elliptic, ovate-elliptic, or ovate-lanceolate, 5–12(–17) × 3–7 mm, base rounded to cuneate-attenuate, margins entire or sparsely dentate to serrate, apex acuminate to acute or obtuse, surfaces villous-glandular, hairs 0.2–0.8 mm, gland-tipped.
Flowers	herkogamous, 1–10, from proximal to distal nodes.	herkogamous, 1–22, from proximal to distal nodes.
Styles	glabrous.	glabrous.
Corollas	yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate; tube-throat funnelform, 7–8 mm, exserted beyond calyx margin; lobes oblong, apex rounded to truncate.	yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 9–12(–14) mm, exserted beyond calyx margin; lobes broadly obovate, apex rounded.
Fruiting pedicels	10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.	divergent-arcuate, 10–23 mm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped.
Fruiting calyces	tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect.	usually red-dotted, narrowly campanulate, not or weakly inflated, 5–7(–9) mm, margins distinctly toothed or lobed, villous-glandular, ribs angled, lobes pronounced, erect.
Capsules	included, 4–6 mm.	included, 4–7 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 32.	= 32.

	Erythranthe patula	Erythranthe arenaria
Phenology	Flowering Apr–May(–Aug).	Flowering May–Sep.
Habitat	Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices.	Sandy flats, bars, gullies, washes, trails, roadcuts, seasonal creek beds and drainages, rocky slopes, seepy loam, ditches, lake edges, meadows, openings in pine-fir and pine-oak woodlands.
Elevation	200–1900(–2900) m. (700–6200(–9500) ft.)	(100–)500–2600(–2800) m. ((300–)1600–8500(–9200) ft.)
Distribution	ID; MT; OR; WA; WY; AB; BC [WildflowerSearch map]	CA [WildflowerSearch map]
Discussion	Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe arenaria is known from a cluster of six counties of the central Sierra Nevada: Fresno, Madera, Mariposa, Merced, Tulare, and Tuolumne. Most plants of Erythranthe arenaria have relatively even-sized cauline leaves, all sessile to proximally subsessile. Plants in the Yosemite area with persistent basal leaves that are short-petiolate, ovate with a cuneate base, and relatively larger than the more distal cauline ones, and possibly related to E. arenaria, have been named M. floribundus var. subulatus. These might be construed as showing the influence of E. geniculata, but that species occurs only at the lower range of elevation of E. arenaria, while plants referable to Mimulus floribundus var. subulatus occur at least to 2300 m and also have the erect habit characteristic of E. arenaria. These variants should be investigated, especially in the Yosemite area where they appear to be relatively common, with the possibility that they indeed represent a distinct entity. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 397.	FNA vol. 17, p. 403.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus patulus	Mimulus arenarius, M. floribundus var. subulatus, M. multiflorus, M. subulatus, M. trisulcatus
Name authority	(Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012)	(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 38. (2012)
Web links	Local floras: BC, OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe arenaria

stalk-leaf monkey-flower

sand-loving monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Stems

erect to ascending, straight or geniculate at nodes, usually simple, (3–)5–15(–24) cm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.

erect to ascending, straight or geniculate at nodes, simple or branched, 5–20 cm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped.

Leaves

cauline, basal not persistent;

petiole (5–)8–25 mm;

blade palmately 3-veined, deltate or ovate to ovate-lanceolate, 4–12(–17) × 3–10(–14) mm, base rounded to cuneate-truncate, margins usually denticulate, apex acute to obtuse, surfaces stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.

basal and cauline;

petiole 0 mm or proximals 1–3(–5) mm;

blade 1-veined or palmately 3-veined, elliptic to narrowly elliptic, ovate-elliptic, or ovate-lanceolate, 5–12(–17) × 3–7 mm, base rounded to cuneate-attenuate, margins entire or sparsely dentate to serrate, apex acuminate to acute or obtuse, surfaces villous-glandular, hairs 0.2–0.8 mm, gland-tipped.

Flowers

herkogamous, 1–10, from proximal to distal nodes.

herkogamous, 1–22, from proximal to distal nodes.

Styles

glabrous.

Corollas

yellow, abaxial limb usually with a few red or brownish dots, radially or bilaterally symmetric, regular or weakly bilabiate;

tube-throat funnelform, 7–8 mm, exserted beyond calyx margin;

lobes oblong, apex rounded to truncate.

yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate;

tube-throat funnelform, 9–12(–14) mm, exserted beyond calyx margin;

lobes broadly obovate, apex rounded.

Fruiting pedicels

10–25(–38) mm, stipitate-glandular, hairs 0.2–0.5 mm, gland-tipped.

divergent-arcuate, 10–23 mm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped.

Fruiting calyces

tubular, weakly or not inflated, 5–6(–7) mm, margins distinctly toothed or lobed, sparsely stipitate-glandular to sparsely hirtellous, lobes pronounced, erect.

usually red-dotted, narrowly campanulate, not or weakly inflated, 5–7(–9) mm, margins distinctly toothed or lobed, villous-glandular, ribs angled, lobes pronounced, erect.

Capsules

included, 4–6 mm.

included, 4–7 mm.

Anthers

included, glabrous.

= 32.

Erythranthe patula

Erythranthe arenaria

Phenology

Flowering Apr–May(–Aug).

Flowering May–Sep.

Habitat

Ephemeral seeps, springs, rocky stream banks, moist basalt, fine gravel on bedrock, muddy hillside seeps, crevices.

Sandy flats, bars, gullies, washes, trails, roadcuts, seasonal creek beds and drainages, rocky slopes, seepy loam, ditches, lake edges, meadows, openings in pine-fir and pine-oak woodlands.

Elevation

200–1900(–2900) m. (700–6200(–9500) ft.)

(100–)500–2600(–2800) m. ((300–)1600–8500(–9200) ft.)

Distribution

ID; MT; OR; WA; WY; AB; BC

[WildflowerSearch map]

Discussion

Erythranthe patula is distinctive with its long-petiolate leaves with ovate blades and its small, weakly bilabiate to nearly radially symmetric corollas. Vestiture may include only minute, stipitate-glandular hairs or it may be an intergrading mix of stipitate-glandular hairs and minute (0.1–0.2 mm), sharp-pointed, eglandular hairs. Plants may have stipitate-glandular pedicels and calyces but hirtellous, eglandular stems, or they may have stipitate-glandular stems and pedicels but hirtellous calyces.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe arenaria is known from a cluster of six counties of the central Sierra Nevada: Fresno, Madera, Mariposa, Merced, Tulare, and Tuolumne.

Most plants of Erythranthe arenaria have relatively even-sized cauline leaves, all sessile to proximally subsessile. Plants in the Yosemite area with persistent basal leaves that are short-petiolate, ovate with a cuneate base, and relatively larger than the more distal cauline ones, and possibly related to E. arenaria, have been named M. floribundus var. subulatus. These might be construed as showing the influence of E. geniculata, but that species occurs only at the lower range of elevation of E. arenaria, while plants referable to Mimulus floribundus var. subulatus occur at least to 2300 m and also have the erect habit characteristic of E. arenaria. These variants should be investigated, especially in the Yosemite area where they appear to be relatively common, with the possibility that they indeed represent a distinct entity.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 397.

FNA vol. 17, p. 403.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus patulus

Mimulus arenarius, M. floribundus var. subulatus, M. multiflorus, M. subulatus, M. trisulcatus

Name authority

(Pennell) G. L. Nesom: Phytoneuron 2012-39: 39. (2012)

(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 38. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Erythranthe patula

Erythranthe arenaria

Erythranthe patula

Erythranthe arenaria