Erythranthe palmeri |
Phrymaceae |
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Palmer's monkeyflower |
lop-seed family, monkeyflower family |
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Habit | Annuals, taprooted. | Herbs, subshrubs, or shrubs, annual or perennial, aquatic or terrestrial, sometimes fleshy, autotrophic. | ||||||||||||||||||||||||
Stems | erect, simple or branched from basal nodes, 4–17 cm, minutely puberulent. |
erect or ascending to prostrate, 4-angled, sometimes winged. |
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Leaves | cauline, basal not persistent; petiole 0 mm; blade pinnately veined, palmately 3-veined (in broader ones), linear to oblanceolate, (3–)4–17 × 1–4 mm, base attenuate, margins entire, apex acute to obtuse, surfaces minutely puberulent. |
deciduous or persistent, basal and cauline or all cauline, rarely subrosulate or rosulate (Erythranthe), opposite, or alternate distally, simple; stipules absent; petiole present or absent; blade fleshy, semi-fleshy, or not, not leathery, margins entire or toothed. |
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Inflorescences | terminal and axillary racemes or flowers solitary (Glossostigma, some Erythranthe, some annual plants); flowers erect to nodding or strongly reflexed and appressed to inflorescence axis (Phryma). |
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Flowers | herkogamous, 1–36, from distal or medial to distal nodes. |
bisexual, perianth and androecium hypogynous; sepals (3 or)4 or 5, proximally connate, calyx radially or bilaterally symmetric; petals 3–5, proximally connate, corolla bilaterally symmetric, rarely nearly radially in reduced forms, strongly to weakly bilabiate, rarely nearly regular, salverform to tubular-funnelform, funnelform, campanulate, or compressed; stamens (2–)4, adnate to corolla, didynamous [both pairs of equal length in autogamous forms], staminode 0; pistil 1, 2-carpellate, ovary superior, (1 or)2-locular, placentation axile, basal (Phryma), or parietal (Diplacus, Mimetanthe); ovules anatropous or orthotropous (Phryma), unitegmic, tenuinucellate; style 1; stigma 1, 2-lobed. |
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Styles | glabrous. |
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Corollas | pink to purple, abaxial limb with 2 yellow ridges, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 6–15 mm, exserted beyond calyx margin; limb expanded 8–15 mm, lobes deeply notched, abaxial limb sparsely bearded. |
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Fruiting pedicels | spreading horizontally, 5–33 mm. |
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Fruiting calyces | sometimes red-spotted on ribs, becoming straw colored, cylindric, 4–8 mm, margins distinctly toothed or lobed, glabrous, ribs weak, lobes pronounced, erect, margins ciliate. |
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Fruits | capsules, dehiscence loculicidal [septicidal or irregular], or achenes [berry]. |
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Capsules | included, 4–8 mm. |
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Seeds | 1–2000, yellowish brown or brown, narrowly ellipsoid, slightly flattened bilaterally; embryo straight, endosperm sparse. |
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Anthers | included, glabrous. |
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Erythranthe palmeri |
Phrymaceae |
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Phenology | Flowering Apr–Jun. | |||||||||||||||||||||||||
Habitat | Moist areas in openings in pine forest and desert chaparral transitions. | |||||||||||||||||||||||||
Elevation | 900–2200 m. (3000–7200 ft.) | |||||||||||||||||||||||||
Distribution |
CA
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North America; Mexico; Central America; w South America (primarily Andean); s Asia (India); se Asia; e Africa; Indian Ocean Islands (Madagascar); Pacific Islands (New Zealand); Australia [Introduced in Europe, s Africa] |
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Discussion | Erythranthe palmeri has been confused with other closely related species, including E. diffusa, E. discolor (pink form), E. rhodopetra, and E. sierrae. It was previously thought to be a widely distributed species because of this taxonomic confusion but now is regarded as endemic to the Transverse Range in the San Bernardino and San Gabriel mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 13, species ca. 200 (6 genera, 139 species in the flora). Over one half of the species in Phrymaceae are members of Diplacus and Erythranthe and together include over 160 species; all other genera each have seven or fewer species. Until recently, Phrymaceae consisted only of Phryma leptostachya, a taxonomically isolated species of eastern North America and eastern Asia. Molecular studies have established a relationship not with the Verbenaceae, as was earlier postulated (see H. L. Whipple 1972; R. Venkata Ramana et al. 2000), but rather with Mimulus and other genera, suggesting that Phrymaceae should be enlarged. The sequence of genera in Phrymaceae here follows the phylogeny proposed by P. M. Beardsley and R. G. Olmstead (2002). One of the major lineages of Phrymaceae is primarily a Southern Hemisphere group ranging from Australia and New Zealand to southeastern and south Asia (India), Madagascar, and South Africa. Mimulus in the narrow sense, including the two endemic North American species, is part of this group, which includes 24 species in seven genera. The largest major lineage includes 158 species in five genera from North America, South America, and southeast Asia. This lineage includes two genera from Mexico and Central America: Hemichaena Bentham, which is sister to the North American Diplacus and Mimetanthe, and Leucocarpus D. Don, which is sister to the American and Asian Erythranthe. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 387. | FNA vol. 17, p. 365. | ||||||||||||||||||||||||
Parent taxa | Phrymaceae > Erythranthe | |||||||||||||||||||||||||
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Synonyms | Mimulus palmeri | |||||||||||||||||||||||||
Name authority | (A. Gray) N. S. Fraga: Phytoneuron 2012-39: 35. (2012) | Schauer | ||||||||||||||||||||||||
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