Erythranthe nudata |
Erythranthe scouleri |
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bare monkeyflower |
Columbia River monkey-flower, Scouler's monkey-flower |
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Habit | Annuals, taprooted or fibrous-rooted. | Perennials, rhizomatous, producing long, sparsely leafy runners from basal nodes. |
Stems | erect or ascending, simple or few-branched from basal nodes, branches mostly reddish purple, (5–)9–30 cm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes. |
erect, simple or few-branched, 15–80 cm, glabrous. |
Leaves | cauline on wide internodes; petiole 5–30 mm, distals 0 mm; blade 1-veined or palmately 3–5-veined, proximals lanceolate or oblong-lanceolate to ovate, narrowly spatulate, or oblanceolate, distals usually linear, not perfoliate, 5–15(–30) × 1–5 mm, base attenuate, margins denticulate to proximally dentate-lobed, apex acute, surfaces glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes. |
cauline, basal usually not persistent; petiole: proximals and medial 10–25 mm (gradually merging into blade), distals 0 mm; blade palmately (3–)5–7-veined to subpinnate, (proximal to medial) oblong-elliptic to oblong-lanceolate, 25–60 × 8–18 mm, usually 3–4 times longer than wide, base attenuate, margins evenly, shallowly dentate or crenate to mucronate or mucronulate, teeth 10–20 per side, sometimes more deeply toothed at base, apex obtuse to acute, surfaces glabrous. |
Flowers | herkogamous, (2–)4–8, usually in proximal or medial to distal axils, chasmogamous. |
herkogamous, (1 or)2–8, from distal nodes. |
Styles | glabrous. |
minutely, prominently hirsutulous to villosulous. |
Corollas | yellow, without a large red splotch, throat floor and tube red-spotted, bilaterally symmetric, bilabiate; tube-throat cylindric-funnelform, 8–12 mm, exserted 2–4 mm beyond calyx margin; limb expanded 8–12 mm. |
yellow, without red markings, bilaterally symmetric, bilabiate; tube-throat funnelform, 20–24 mm, exserted 10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
Fruiting pedicels | erect in flower, spreading to divaricate, rarely recurved, in fruit, 10–35 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes. |
20–25 mm, glabrous. |
Fruiting calyces | ovate-campanulate, inflated, sagittally compressed, 6–13 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes, throat closing, abaxial lobe upcurving over lateral ones, nearly closing orifice. |
ovoid, inflated, sagittally compressed, 13–14 mm, glabrous, throat closing. |
Capsules | included, 6–7 mm. |
unknown. |
Anthers | included, glabrous. |
included, glabrous. |
Erythranthe nudata |
Erythranthe scouleri |
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Phenology | Flowering Apr–Jun. | Flowering May–Aug. |
Habitat | Open gravelly seeps on serpentine outcrops, serpentine crevices, springs, streamsides, gravelly creek beds, roadside drainages and swales. | Salt marshes, river banks. |
Elevation | 200–700 m. (700–2300 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
CA
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OR; WA |
Discussion | Erythranthe nudata is distinct in its annual duration, few, inconspicuous, and narrow leaves, long and spreading-divaricate pedicels, and large corollas. The plants apparently are restricted to serpentine substrate and known only from Colusa, Glenn, Lake, Mendocino, Napa, and Sonoma counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe scouleri is distinctive in its oblong-elliptic leaves with long-tapering bases and closely toothed margins, completely glabrous vestiture, long, leafy runners from basal cauline nodes, large corollas with broad limbs, and prominently hairy styles. Several features suggest a close relationship to E. decora, particularly its very large corollas, hairy styles, closely toothed leaf margins, tall, simple, and erect stems, numerous runners, and its geographic range. All collections apparently have been made near the mouth of the Columbia River in Clatsop and Columbia counties, Oregon (G. L. Nesom 2013d). Recent observations (Alexander John Wright, pers. comm.; photos http://www.inaturalist.org/observations/3537008) indicate that it also occurs in Wahkiakum County, Washington, in the Julia Butler Hansen Refuge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 417. | FNA vol. 17, p. 415. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus nudatus | Mimulus scouleri, M. guttatus subsp. scouleri |
Name authority | (Curran ex Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Hooker) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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