FNA: Erythranthe nudata vs. Erythranthe hymenophylla

	Erythranthe nudata	Erythranthe hymenophylla
	bare monkeyflower	thin-sepal monkeyflower
Habit	Annuals, taprooted or fibrous-rooted.	Annuals, filiform-taprooted.
Stems	erect or ascending, simple or few-branched from basal nodes, branches mostly reddish purple, (5–)9–30 cm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes.	prostrate to ascending-erect, sharply bent at basal nodes, simple or few-branched, 5–25 cm, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped.
Leaves	cauline on wide internodes; petiole 5–30 mm, distals 0 mm; blade 1-veined or palmately 3–5-veined, proximals lanceolate or oblong-lanceolate to ovate, narrowly spatulate, or oblanceolate, distals usually linear, not perfoliate, 5–15(–30) × 1–5 mm, base attenuate, margins denticulate to proximally dentate-lobed, apex acute, surfaces glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes.	basal and cauline, largest at mid stem; petiole 6–30 mm; blade pinnately veined, broadly lanceolate to ovate, 10–35 × 10–30 mm, distinctly membranous, base cuneate to shallowly cordate, margins coarsely dentate to shallowly denticulate or entire, apex acute to obtuse, surfaces glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped, glandular.
Flowers	herkogamous, (2–)4–8, usually in proximal or medial to distal axils, chasmogamous.	herkogamous, 1–6, from proximal to distal nodes.
Styles	glabrous.	glabrous.
Corollas	yellow, without a large red splotch, throat floor and tube red-spotted, bilaterally symmetric, bilabiate; tube-throat cylindric-funnelform, 8–12 mm, exserted 2–4 mm beyond calyx margin; limb expanded 8–12 mm.	light yellow, throat and abaxial lobes red- or purple-spotted, sometimes with small white patches, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 10–14 mm, exserted beyond calyx margin; lobes obovate-oblong, apex rounded to truncate or notched.
Fruiting pedicels	erect in flower, spreading to divaricate, rarely recurved, in fruit, 10–35 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes.	divergent at right angles from stem, usually closely paired, 10–45 mm, negatively phototropic, causing capsules to be pressed against a cliff face or crevice at time of dehiscence, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped.
Fruiting calyces	ovate-campanulate, inflated, sagittally compressed, 6–13 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes, throat closing, abaxial lobe upcurving over lateral ones, nearly closing orifice.	angled, tubular-campanulate, slightly inflated, 5–7 mm, margins distinctly toothed or lobed, sparsely stipitate-glandular, lobes pronounced, erect.
Capsules	included, 6–7 mm.	included, 3–6 mm.
Anthers	included, glabrous.	included, glabrous.
2n		= 32.

	Erythranthe nudata	Erythranthe hymenophylla
Phenology	Flowering Apr–Jun.	Flowering Apr–Aug(–Sep).
Habitat	Open gravelly seeps on serpentine outcrops, serpentine crevices, springs, streamsides, gravelly creek beds, roadside drainages and swales.	Steep, seasonally moist, basalt cliffs with west or southwest exposure, mesic coniferous forests.
Elevation	200–700 m. (700–2300 ft.)	800–1300 m. (2600–4300 ft.)
Distribution	CA [WildflowerSearch map]	ID; MT; OR
Discussion	Erythranthe nudata is distinct in its annual duration, few, inconspicuous, and narrow leaves, long and spreading-divaricate pedicels, and large corollas. The plants apparently are restricted to serpentine substrate and known only from Colusa, Glenn, Lake, Mendocino, Napa, and Sonoma counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)	In the protologue, R. J. Meinke observed that plants of Erythranthe hymenophylla have reflexed fruiting pedicels that increase seed dispersal back onto the vertical cliff wall, the characteristic habitat of the species. The hanging habit of E. hymenophylla is reflected in a sharp (90º to 180º) bend in the basal nodes and the long pedicels that are closely paired and divergent in parallel at about right angles from the stem. The species also is characterized by it very short calyx to corolla length, relatively short capsules, and large seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 417.	FNA vol. 17, p. 397.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus nudatus	Mimulus hymenophyllus
Name authority	(Curran ex Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012)	(Meinke) G. L. Nesom: Phytoneuron 2012-39: 38. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe hymenophylla

bare monkeyflower

thin-sepal monkeyflower

Habit

Annuals, taprooted or fibrous-rooted.

Annuals, filiform-taprooted.

Stems

erect or ascending, simple or few-branched from basal nodes, branches mostly reddish purple, (5–)9–30 cm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes.

prostrate to ascending-erect, sharply bent at basal nodes, simple or few-branched, 5–25 cm, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped.

Leaves

cauline on wide internodes;

petiole 5–30 mm, distals 0 mm;

blade 1-veined or palmately 3–5-veined, proximals lanceolate or oblong-lanceolate to ovate, narrowly spatulate, or oblanceolate, distals usually linear, not perfoliate, 5–15(–30) × 1–5 mm, base attenuate, margins denticulate to proximally dentate-lobed, apex acute, surfaces glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes.

basal and cauline, largest at mid stem;

petiole 6–30 mm;

blade pinnately veined, broadly lanceolate to ovate, 10–35 × 10–30 mm, distinctly membranous, base cuneate to shallowly cordate, margins coarsely dentate to shallowly denticulate or entire, apex acute to obtuse, surfaces glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped, glandular.

Flowers

herkogamous, (2–)4–8, usually in proximal or medial to distal axils, chasmogamous.

herkogamous, 1–6, from proximal to distal nodes.

Styles

glabrous.

Corollas

yellow, without a large red splotch, throat floor and tube red-spotted, bilaterally symmetric, bilabiate;

tube-throat cylindric-funnelform, 8–12 mm, exserted 2–4 mm beyond calyx margin;

limb expanded 8–12 mm.

light yellow, throat and abaxial lobes red- or purple-spotted, sometimes with small white patches, bilaterally symmetric, weakly bilabiate;

tube-throat funnelform, 10–14 mm, exserted beyond calyx margin;

lobes obovate-oblong, apex rounded to truncate or notched.

Fruiting pedicels

erect in flower, spreading to divaricate, rarely recurved, in fruit, 10–35 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes.

divergent at right angles from stem, usually closely paired, 10–45 mm, negatively phototropic, causing capsules to be pressed against a cliff face or crevice at time of dehiscence, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped.

Fruiting calyces

ovate-campanulate, inflated, sagittally compressed, 6–13 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes, throat closing, abaxial lobe upcurving over lateral ones, nearly closing orifice.

angled, tubular-campanulate, slightly inflated, 5–7 mm, margins distinctly toothed or lobed, sparsely stipitate-glandular, lobes pronounced, erect.

Capsules

included, 6–7 mm.

included, 3–6 mm.

Anthers

included, glabrous.

= 32.

Erythranthe nudata

Erythranthe hymenophylla

Phenology

Flowering Apr–Jun.

Flowering Apr–Aug(–Sep).

Habitat

Open gravelly seeps on serpentine outcrops, serpentine crevices, springs, streamsides, gravelly creek beds, roadside drainages and swales.

Steep, seasonally moist, basalt cliffs with west or southwest exposure, mesic coniferous forests.

Elevation

200–700 m. (700–2300 ft.)

800–1300 m. (2600–4300 ft.)

Distribution

[WildflowerSearch map]

ID; MT; OR

Discussion

Erythranthe nudata is distinct in its annual duration, few, inconspicuous, and narrow leaves, long and spreading-divaricate pedicels, and large corollas. The plants apparently are restricted to serpentine substrate and known only from Colusa, Glenn, Lake, Mendocino, Napa, and Sonoma counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In the protologue, R. J. Meinke observed that plants of Erythranthe hymenophylla have reflexed fruiting pedicels that increase seed dispersal back onto the vertical cliff wall, the characteristic habitat of the species. The hanging habit of E. hymenophylla is reflected in a sharp (90º to 180º) bend in the basal nodes and the long pedicels that are closely paired and divergent in parallel at about right angles from the stem. The species also is characterized by it very short calyx to corolla length, relatively short capsules, and large seeds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 417.

FNA vol. 17, p. 397.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus nudatus

Mimulus hymenophyllus

Name authority

(Curran ex Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012)

(Meinke) G. L. Nesom: Phytoneuron 2012-39: 38. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Erythranthe nudata

Erythranthe hymenophylla

Erythranthe nudata

Erythranthe hymenophylla