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bare monkeyflower

large monkey-flower, magnificent monkeyflower, magnificent seep monkeyflower

Habit Annuals, taprooted or fibrous-rooted. Perennials, rhizomatous, sometimes rooting at proximal nodes.
Stems

erect or ascending, simple or few-branched from basal nodes, branches mostly reddish purple, (5–)9–30 cm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes.

erect, sometimes decumbent basally, branched, often fistulose, (25–)50–120(–160) cm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs.

Leaves

cauline on wide internodes;

petiole 5–30 mm, distals 0 mm;

blade 1-veined or palmately 3–5-veined, proximals lanceolate or oblong-lanceolate to ovate, narrowly spatulate, or oblanceolate, distals usually linear, not perfoliate, 5–15(–30) × 1–5 mm, base attenuate, margins denticulate to proximally dentate-lobed, apex acute, surfaces glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes.

basal and cauline, basal usually not persistent, bracteate in inflorescence;

petiole 10–80 mm, gradually reduced distally;

blade subpinnately, sometimes palmately, 5–7-veined, ovate to broadly elliptic, 25–60 × 20–40(–60) mm, usually 1–2 times longer than wide, base truncate or truncate-cuneate to subcordate, margins crenulate to dentate, proximally sometimes sublyrate, apex rounded to obtuse, surfaces of distals densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs.

Flowers

herkogamous, (2–)4–8, usually in proximal or medial to distal axils, chasmogamous.

herkogamous, 8–26, mostly from distal nodes, usually in bracteate racemes.

Styles

glabrous.

hirtellous.

Corollas

yellow, without a large red splotch, throat floor and tube red-spotted, bilaterally symmetric, bilabiate;

tube-throat cylindric-funnelform, 8–12 mm, exserted 2–4 mm beyond calyx margin;

limb expanded 8–12 mm.

yellow, red-dotted within, bilaterally symmetric, bilabiate;

tube-throat broadly funnelform, (14–)16–24 mm, exserted (8–)10–15 mm beyond calyx margin;

limb broadly expanded.

Fruiting pedicels

erect in flower, spreading to divaricate, rarely recurved, in fruit, 10–35 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes.

10–35 mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs.

Fruiting calyces

ovate-campanulate, inflated, sagittally compressed, 6–13 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes, throat closing, abaxial lobe upcurving over lateral ones, nearly closing orifice.

straight-erect or nodding 45–100º, ovate-campanulate, inflated, sagittally compressed, 15–22(–25) mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs, throat closing.

Capsules

included, 6–7 mm.

included, 8–12 mm.

Anthers

included, glabrous.

included, glabrous.

2n

= 28.

Erythranthe nudata

Erythranthe grandis

Phenology Flowering Apr–Jun. Flowering (Apr–)May–Jul(–Sep).
Habitat Open gravelly seeps on serpentine outcrops, serpentine crevices, springs, streamsides, gravelly creek beds, roadside drainages and swales. Beaches, dunes, coastal bluffs, wet cliff faces, mud flats and seeps, marshes, drainage ditches, creeks, rarely in coastal sage scrub.
Elevation 200–700 m. (700–2300 ft.) 0–200(–800) m. (0–700(–2600) ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CA; OR
[WildflowerSearch map]
Discussion

Erythranthe nudata is distinct in its annual duration, few, inconspicuous, and narrow leaves, long and spreading-divaricate pedicels, and large corollas. The plants apparently are restricted to serpentine substrate and known only from Colusa, Glenn, Lake, Mendocino, Napa, and Sonoma counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The densely, evenly puberulent vestiture of pedicels, calyces, and distal stems usually is diagnostic, especially in combination with the large flowers (corollas and mature calyces) and tall stature. Plants from scattered collections are much shorter than normal but have large corollas and characteristic vestiture.

Erythranthe grandis characteristically occurs in coastal localities from southern California to northern Oregon but also is found in inland localities and habitats near the coast but well away from salt spray.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 417. FNA vol. 17, p. 412.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus nudatus Mimulus guttatus var. grandis, M. grandis, M. guttatus subsp. litoralis, M. langsdorffii var. grandis, M. procerus
Name authority (Curran ex Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)
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