Erythranthe nudata |
Erythranthe arvensis |
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bare monkeyflower |
field monkey-flower, villous-bract monkeyflower, western monkey-flower |
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Habit | Annuals, taprooted or fibrous-rooted. | Annuals, taprooted or fibrous-rooted, sometimes rooting at proximal cauline nodes if decumbent. |
Stems | erect or ascending, simple or few-branched from basal nodes, branches mostly reddish purple, (5–)9–30 cm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes. |
erect to decumbent-ascending, simple or branched from proximal to medial nodes, usually 4-angled, fistulose to very narrow, 5–70 cm, glabrous, sometimes minutely hirtellous in inflorescence, hairs deflexed, eglandular. |
Leaves | cauline on wide internodes; petiole 5–30 mm, distals 0 mm; blade 1-veined or palmately 3–5-veined, proximals lanceolate or oblong-lanceolate to ovate, narrowly spatulate, or oblanceolate, distals usually linear, not perfoliate, 5–15(–30) × 1–5 mm, base attenuate, margins denticulate to proximally dentate-lobed, apex acute, surfaces glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes. |
basal and cauline or basal not persistent, often largest at mid stem or above, reduced in size distally; petiole 3–20(–90) mm, distals 0 mm; blade palmately 3–5-veined, ovate to orbicular, orbicular-ovate, oblong-ovate, or (middle and distal cauline) broadly orbicular to depressed-ovate or nearly reniform, (5–)10–35(–45) × 6–26(–50) mm, distal closely paired, auriculate-subclasping, base rounded to truncate, subcordate, or shallowly cordate, margins denticulate or subentire to distinctly dentate, on larger plants proximal characteristically lacerate-lobed to pinnatifid at margin base, apex rounded, surfaces glabrous except for bracts densely villous abaxially, sometimes also adaxially, hairs long, sometimes vitreous, flattened, eglandular, multicellular. |
Flowers | herkogamous, (2–)4–8, usually in proximal or medial to distal axils, chasmogamous. |
plesiogamous, 3–8(–16), from remote distal nodes, chasmogamous or cleistogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, without a large red splotch, throat floor and tube red-spotted, bilaterally symmetric, bilabiate; tube-throat cylindric-funnelform, 8–12 mm, exserted 2–4 mm beyond calyx margin; limb expanded 8–12 mm. |
yellow, usually red-spotted, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat cylindric-funnelform, (7–)8–12 mm, exserted (0–)1–2(–3) mm beyond calyx margin; limb expanded 5–10 mm. |
Fruiting pedicels | erect in flower, spreading to divaricate, rarely recurved, in fruit, 10–35 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes. |
5–40(–90) mm, longer than subtending leaves, glabrous. |
Fruiting calyces | ovate-campanulate, inflated, sagittally compressed, 6–13 mm, glabrous or minutely stipitate-glandular, hairs 0.05–0.1 mm, at least just above nodes, throat closing, abaxial lobe upcurving over lateral ones, nearly closing orifice. |
red-dotted or not, ovate-campanulate, inflated, sagittally compressed, (7–)9–14 mm, minutely hirtellous, throat closing or not, remaining open, lobes upcurving weakly, adaxial lobe not distinctly longer than abaxial, not falcate. |
Capsules | included, 6–7 mm. |
included, stipitate, (5–)6–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
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Erythranthe nudata |
Erythranthe arvensis |
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Phenology | Flowering Apr–Jun. | Flowering Apr–Jun(–Jul). |
Habitat | Open gravelly seeps on serpentine outcrops, serpentine crevices, springs, streamsides, gravelly creek beds, roadside drainages and swales. | Hills, ridges, clay banks, stream banks, moist woods. |
Elevation | 200–700 m. (700–2300 ft.) | 30–1900(–2300) m. (100–6200(–7500) ft.) |
Distribution |
CA
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CA; ID; MT; NV; OR; UT; WA; WY; BC; Mexico (Baja California)
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Discussion | Erythranthe nudata is distinct in its annual duration, few, inconspicuous, and narrow leaves, long and spreading-divaricate pedicels, and large corollas. The plants apparently are restricted to serpentine substrate and known only from Colusa, Glenn, Lake, Mendocino, Napa, and Sonoma counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe arvensis usually is easily recognized, characterized by its annual duration (but commonly rooting at proximal cauline nodes, suggestive of a rhizomatous habit), glabrous stems with nodes relatively few and remotely spaced, depressed-ovate leaves with margins often sublyrate (lacerate-lobed to subpinnatifid) at the base, distal leaves and bracts densely villous with vitreous eglandular hairs, other leaves glabrous, and corollas varying in size from relatively small but perhaps chasmogamous (the type of Mimulus arvensis) to even smaller (cleistogamous; the type of M. micranthus). The breeding system is consistently autogamous. The relatively short and even-sized calyx lobes that do not turn upward to close the orifice have been considered diagnostic of E. arvensis. This feature is evident in some plants, but others (perhaps reflecting gene flow from other species) have a longer adaxial calyx lobe and abaxial lobes that turn upward variably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 417. | FNA vol. 17, p. 420. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus nudatus | Mimulus arvensis, M. guttatus subsp. arvensis, M. guttatus var. arvensis, M. guttatus subsp. micranthus, M. langsdorffii var. arvensis, M. longulus, M. micranthus |
Name authority | (Curran ex Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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