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Kaweah monkeyflower, Norris' monkeyflower

field monkey-flower, villous-bract monkeyflower, western monkey-flower

Habit Annuals, fibrous-rooted or filiform-taprooted. Annuals, taprooted or fibrous-rooted, sometimes rooting at proximal cauline nodes if decumbent.
Stems

ascending to erect-ascending, geniculate at nodes, usually branched from proximal nodes, 2–15(–25) cm, villous-glandular.

erect to decumbent-ascending, simple or branched from proximal to medial nodes, usually 4-angled, fistulose to very narrow, 5–70 cm, glabrous, sometimes minutely hirtellous in inflorescence, hairs deflexed, eglandular.

Leaves

basal and cauline;

petiole 5–10(–15) mm;

blade palmately 3–5-veined, sometimes with 1–3 distal vein pairs diverging pinnately, elliptic to elliptic-obovate, 20–35 × 10–20 mm, base usually attenuate, margins subentire to distally denticulate, apex acute to obtuse, surfaces villous-glandular.

basal and cauline or basal not persistent, often largest at mid stem or above, reduced in size distally;

petiole 3–20(–90) mm, distals 0 mm;

blade palmately 3–5-veined, ovate to orbicular, orbicular-ovate, oblong-ovate, or (middle and distal cauline) broadly orbicular to depressed-ovate or nearly reniform, (5–)10–35(–45) × 6–26(–50) mm, distal closely paired, auriculate-subclasping, base rounded to truncate, subcordate, or shallowly cordate, margins denticulate or subentire to distinctly dentate, on larger plants proximal characteristically lacerate-lobed to pinnatifid at margin base, apex rounded, surfaces glabrous except for bracts densely villous abaxially, sometimes also adaxially, hairs long, sometimes vitreous, flattened, eglandular, multicellular.

Flowers

herkogamous, 1–5, from medial to distal nodes.

plesiogamous, 3–8(–16), from remote distal nodes, chasmogamous or cleistogamous.

Styles

glabrous.

glabrous.

Corollas

yellow, base of each lobe with a prominent maroon splotch, abaxial limb with white patch at 2 sinus bases, weakly bilaterally or radially symmetric, weakly bilabiate or regular;

tube-throat cylindric-funnelform, 12–16 mm, exserted beyond calyx margin;

limb expanded 15–30 mm, lobes oblong-obovate to orbicular-obovate, apex rounded-truncate.

yellow, usually red-spotted, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular;

tube-throat cylindric-funnelform, (7–)8–12 mm, exserted (0–)1–2(–3) mm beyond calyx margin;

limb expanded 5–10 mm.

Fruiting pedicels

20–35(–50) mm, villous-glandular.

5–40(–90) mm, longer than subtending leaves, glabrous.

Fruiting calyces

red-dotted, campanulate, weakly inflated, 4–6 mm, margins distinctly toothed or lobed, villous-glandular, ribs rounded-thickened, lobes pronounced, erect, often incurved, linear-oblong to oblong-lanceolate, apex rounded to blunt.

red-dotted or not, ovate-campanulate, inflated, sagittally compressed, (7–)9–14 mm, minutely hirtellous, throat closing or not, remaining open, lobes upcurving weakly, adaxial lobe not distinctly longer than abaxial, not falcate.

Capsules

usually slightly exserted, 4–6 mm.

included, stipitate, (5–)6–7 mm.

Anthers

included, glabrous.

included, glabrous.

2n

= 32.

= 28.

Erythranthe norrisii

Erythranthe arvensis

Phenology Flowering Mar–May. Flowering Apr–Jun(–Jul).
Habitat Steep marble outcrops in soil pockets, moss covered marble and quartzite ledges, cracks, fractures, weathered faces, chamise chaparral or blue oak woodlands. Hills, ridges, clay banks, stream banks, moist woods.
Elevation 300–1300 m. (1000–4300 ft.) 30–1900(–2300) m. (100–6200(–7500) ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CA; ID; MT; NV; OR; UT; WA; WY; BC; Mexico (Baja California)
[WildflowerSearch map]
Discussion

Erythranthe norrisii is known only from the Kaweah River drainage; most populations are in Sequoia National Park in Tulare County. The species is characterized by its short-petiolate leaves with attenuate bases, very large corollas with red splotches at the base of each lobe and two white patches on the abaxial limb, and very short, purple-dotted calyces with rounded-thickened ribs and linear-oblong lobes incurved in fruit. The capsules often are slightly exserted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe arvensis usually is easily recognized, characterized by its annual duration (but commonly rooting at proximal cauline nodes, suggestive of a rhizomatous habit), glabrous stems with nodes relatively few and remotely spaced, depressed-ovate leaves with margins often sublyrate (lacerate-lobed to subpinnatifid) at the base, distal leaves and bracts densely villous with vitreous eglandular hairs, other leaves glabrous, and corollas varying in size from relatively small but perhaps chasmogamous (the type of Mimulus arvensis) to even smaller (cleistogamous; the type of M. micranthus). The breeding system is consistently autogamous. The relatively short and even-sized calyx lobes that do not turn upward to close the orifice have been considered diagnostic of E. arvensis. This feature is evident in some plants, but others (perhaps reflecting gene flow from other species) have a longer adaxial calyx lobe and abaxial lobes that turn upward variably.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 405. FNA vol. 17, p. 420.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus norrisii Mimulus arvensis, M. guttatus subsp. arvensis, M. guttatus var. arvensis, M. guttatus subsp. micranthus, M. langsdorffii var. arvensis, M. longulus, M. micranthus
Name authority (Heckard & Shevock) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)
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